Journal of Threatened
Taxa | www.threatenedtaxa.org | 26 February 2025 | 17(2): 26566–26570
ISSN 0974-7907 (Online)
| ISSN 0974-7893 (Print)
https://doi.org/10.11609/jott.9271.17.2.26566-26570
#9271 | Received 05 July
2024 | Final received 09 January 2025 | Finally accepted 01 February 2025
Cuphea carthagenensis (Jacq.) J.F.Macbr. (Lythraceae)
— a new non-native plant record for the Eastern Ghats of India
Prabhat Kumar Das
1, Bishal Kumar Majhi 2, Shashi
Sourav Hansda 3, Samarendra
Narayan Mallick 4, Purnendu Panda 5
& Pratap Chandra Panda 6
1,6
Centre for Biotechnology, Siksha
‘O’ Anusandhan (Deemed to be University), Kalinga
Nagar, Ghatikia, Bhubaneswar, Odisha 751003, India.
2
G.B. Pant National Institute of Himalayan
Environment, Northeast Regional Centre, Chandranagar,
Itanagar, Arunachal Pradesh 791113, India.
3 Department of Botany,
Utkal University, Bhubaneswar, Odisha 751004, India.
4 Department of Botany,
Ravenshaw University, Cuttack, Odisha 753003, India.
5 Central Ayurveda
Research Institute, Bharatpur, Bhubaneswar, Odisha
751029, India.
1 prabhatdasnou@gmail.com,
2 bishalkm@gmail.com, 3 shashibabu087@gmail.com, 4 samarendra.mallick1@yahoo.com,
5 pandapurnendu02@gmail.com,
6 pcpanda2001@yahoo.co.in (corresponding author)
Editor: V. Sampath Kumar, Botanical Survey of India,
Coimbatore, India. Date of publication: 26 February
2025 (online & print)
Citation: Das, P.K., B.K. Majhi, S.S. Hansda, S.N.
Mallick, P. Panda & P.C. Panda (2025). Cuphea carthagenensis (Jacq.) J.F.Macbr. (Lythraceae)
— a new non-native plant record for the Eastern Ghats of India. Journal of Threatened Taxa 17(2): 26566–26570. https://doi.org/10.11609/jott.9271.17.2.26566-26570
Copyright: © Das et al. 2025. Creative Commons Attribution 4.0 International License. JoTT allows unrestricted use, reproduction, and
distribution of this article in any medium by providing adequate credit to the
author(s) and the source of publication.
Funding: None.
Competing interests: The authors declare no competing interests.
Acknowledgements: The authors thank the dean, the School of Pharmaceutical Sciences, and the President, Siksha O Anusandhan Deemed to be University, Bhubaneswar for providing laboratory facilities and encouragement.
During the ongoing
plant diversity inventory of the Eastern Ghat region
of Odisha, the authors collected specimens from two populations of an
interesting species of Cuphea with glandular-pubescent aerial parts and
pale-purple to deep pink flowers from Sunabeda
Plateau and Pendajam waterfall of Koraput District of
Odisha. After examination of plant specimens, consultation of relevant
literature, and matching with authentic herbarium specimens, it is identified
as Cuphea carthagenensis (Jacq.) J.F.Macbr. (Lythraceae),
a species native to South America.
In India,
half-a-dozen species of Cuphea P.Browne
have so far been reported, such as C. hyssopifolia
Kunth, C. carthagenensis
(Jacq.) J.F.Macbr.,
C. ignea A.DC., C. lanceolata
W.T.Aiton, C. micropetala
Kunth, and C. procumbens
Ortega, which are commonly grown as garden ornamental plants or found as
alien weeds in waste places, roadsides and degraded forest areas (Das & Chanda
1987; Singh et al. 2000; Paul & Kumar 2023).
Among these, the
species C. carthagenensis is so far been
reported from northeastern states such as Assam, Arunachal Pradesh, Nagaland (Naithani 1990; Naithani &
Bennet 1990), Mizoram (Singh et al. 2002), Darjeeling Hills of West Bengal
(Paul & Kumar 2012), and Manipur (Panmei et al.
2019) as a naturalized weed in cultivated lands, disturbed habitats and also
sometimes in open forests. There is no report of its occurrence in the Eastern
Ghats of India, which covers the states of Odisha, Andhra Pradesh, Tamil Nadu,
and Karnataka nor from peninsular India. The present report of its occurrence
from Odisha is very interesting in view of its naturalization extending to the
eastern parts of the country (Image 1).
Cuphea carthagenensis (Jacq.) J.F.Macbr.,Publ. Field Columb. Mus., Bot. Ser. 8: 124. 1930; Naithani,
Fl. Pl. India, Nepal & Bhutan: 172. 1990; Naithani
& Bennet, Ind. For. 116(5): 423. 1990; Chowdhery
& al. in P.K. Hajra & al., Mater. Fl. Arunachal Pradesh 1: 496. 1996; Paul
& Kumar, Pleione 6(1): 244. 2012; Graham, Syst. Bot. 42(4): 876. 2017; T.
Mathew & al. in A.A. Mao & S.S. Dash, Fl. Pl. India Annot.
Checkl. Dicot 1: 558. 2020; Paul & Kumar, Fl.
Mahananda Wildlife Sanctuary: 69. 2023. Lythrum
carthagenense Jacq., Enum.
Syst. Pl. 22. 1760. Cuphea pinto (Vand.) Kuntze, Revis. Gen. Pl. 3(3): 96.
1898. Cuphea balsamona Cham. & Schlecht. in Linnaea 2: 363. 1827. Cuphea prunellifolia, FI. Bras. Mer. 3: 108. 1833. (Images 2
& 3)
Common name:
Colombian Cuphea, Colombian wax-weed.
Perennial herb, erect or spreading, much branched from
near the base, 10–40
cm tall, young parts hairy. Stem viscid-pubescent, with
intermixed glandular and non-glandular hairs; trichomes sparse to abundant,
erect, reddish-purple. Leaves opposite, spreading, subsessile
to shortly petiolate, gradually reduced in size in inflorescence; broadly to
narrowly elliptic or elliptic-oblong, rarely ovate, 0.9–3.2 X 0.6–2.0 cm, base
acute to attenuate, apex acute or acuminate, margin entire; upper surface
green, lower surface pale or both surfaces with scattered trichomes, midvein
very prominent, secondary veins 5–6, prominent on dorsal side. Flowers extra
axillary, solitary or in an indistinct leafy raceme, 4–6 mm long; pedicels 1–2
mm long; bracteoles lanceolate-linear. Floral tube, including a descending
spur, sparsely pubescent with glandular hairs, distally rose-purple, otherwise
green, ampullaceous with neck and mouth contracted in
fruit. Calyx tubular, 3–5 mm long, longitudinally ribbed with a short spur at
the base of posterior side; lobes short, equal, deltate,
apex acute or short bristle-tipped. Petals 6, light pink to pink-purple,
subequal, oblong, elliptic, or sub-spatulate, up to 3 mm long. Stamens 11,
slightly longer than the floral tube; 2 short dorsal most inserted deepest,
other 9 alternately unequal in length, inserted at the same level; filaments
cream coloured, anthers light yellow. Ovary oblong,
0.5 mm long, glabrous; ovules 4–6; stigma capitate. Capsules enclosed by
hypanthium, dehiscing on one side; seeds brownish, 2–6, 1–1.5 mm across,
obovate to elliptic or subcircular, apex obtuse to
shallowly retuse, margin thin, flattened, especially
at micropylar end, surface verrucose.
Flowering and
fruiting: February–May.
Distribution: Though
the native range of Cuphea carthagenensis is
South America covering Argentina, Bolivia, Brazil, Colombia, Ecuador, French
Guiana, Guyana, Paraguay, Peru, Suriname, and Venezuela (Graham 2017), it has
widely naturalized outside its native range, in the subtropical and tropical
parts Central America, North America, the Caribbean, Oceania, and Asia. In
Asia, the species is reported to occur in China, East Timor, India, Indonesia,
Japan, Malaysia, Myanmar, Philippines, Singapore, and Taiwan. In Oceania, C.
carthagenensis is known from American Samoa,
Australia, Micronesia, Fiji, French Polynesia, New Caledonia, Papua New Guinea,
Western Samoa, Tonga, and Vanuatu. While it is a dominant weed of rice fields
in Assam, India; it invades corn (Zea mays)
plantings in Indonesia, Coconut Cocos nucifera groves in Vanuatu and
Taro Colocasia esculenta fields in Fiji
(Bradley 2022).
In India, the species
has been previously reported from northeastern states (Assam, Arunachal
Pradesh, Mizoram, Meghalaya, Nagaland, Manipur), higher hills of Darjeeling in
West Bengal, and now from Koraput, Odisha located in the eastern region of the
country indicating its spread as an invasive species.
Specimens examined:
2544/CBT, 20.iv.2024, India, Odisha, Koraput, Sunabeda,
Central University of Odisha campus, 18.701 N & 83.012 E, 2,112 m, coll.
P.K. Das & P.C. Panda; 2545/CBT, 14.v.2024, India, Odisha, Koraput, Pendajam waterfall, on the way to Putsil,
18.739 N & 82.808 E, 1,976 m, coll. P.K. Das and S.N. Mallick.
Specimen images viewed
online: 1650 (K), 600 m, coll. M. Etuge, Mt. Kupe, Cameroon
(K000026893); 9500 (K), 2,000 m, coll. M. Cheek, Muanenguba,
Cameroon (K000026892); 9559 (K), 1,200 m, coll. M. Cheek, Muambong
to Mwabag, Cameroon (K000026894).
For
images - - click here for full PDF
References
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