Journal of Threatened
Taxa | www.threatenedtaxa.org | 26 January 2025 | 17(1): 26353–26369
ISSN 0974-7907 (Online)
| ISSN 0974-7893 (Print)
https://doi.org/10.11609/jott.9210.17.1.26353-26369
#9210 | Received 14 June
2024 | Final received 29 November 2024 | Finally accepted 03 January 2025
Checklist
of forensically significant Rove beetles (Coleoptera: Staphylinidae:
Staphylininae: Staphylinini) from India
Meenakshi Bharti
1 & Shweta Sharma 2
1,2 Department of Zoology
and Environmental Sciences, Punjabi University, Patiala, Punjab 147002, India.
1 adubharti@gmail.com
(corresponding author), 2 shweta.8.6.2023@gmail.com
Editor: Shiju T. Raj, St. Joseph’s College,
Kozhikode, India. Date of publication: 26 January 2025
(online & print)
Citation: Bharti, M. & S. Sharma (2025). Checklist of forensically significant Rove beetles (Coleoptera:
Staphylinidae: Staphylininae: Staphylinini) from India. Journal
of Threatened Taxa 17(1): 26353–26369. https://doi.org/10.11609/jott.9210.17.1.26353-26369
Copyright: © Bharti & Sharma 2025. Creative Commons Attribution 4.0 International License. JoTT allows
unrestricted use, reproduction, and distribution of this article in any medium
by providing adequate credit to the author(s) and the source of publication.
Funding: Financial assistance rendered by Department of Science and Technology,
Ministry of Science and Technology, New Delhi vide grant no.
SR/WOS-A/LS-187/2019 and IF230060 is gratefully acknowledged.
Competing interests: The authors declare no competing interests.
Author details: Dr. Meenakshi Bharti is currently based at the Department of Zoology and Environmental
Sciences Punjabi University, Patiala. Her prime area of interest is forensic
entomology, insect systematics, chemical ecology
and molecular phylogeny. She is one of the leading experts in the field of
forensic entomology in India and is trying her best to popularise this applied
field in the country. She is the first one to generate insect data, which has the potential to be applicable in solving criminal
cases. Besides, she aims to streamline the taxonomy of dipteran flies from
India with major focus on family Calliphoridae
and is also actively involved in such studies from other countries. Her other significant contributions are in the field of ant ecology
and molecular phylogeny of blow flies. Ms. Shweta Sharma, is pursuing her PhD in the
department of Zoology and Environmental Sciences, Punjabi University, Patiala.
Her research focuses on the study of antennal sensilla in
beetles particularly in relation to their niche selection.
Author contributions: The authors contributed equally in
the preparation of the manuscript.
Abstract: Beetles play a
substantial role in calculating the postmortem interval (PMI) during the later
stages of decomposition of a carcass. The dominant families are Silphidae, Dermestidae,
Histeridae, Tenebrionidae, Trogidae, and Staphylinidae. For accuracy in PMI
calculations based on insect fauna, it becomes imperative to catalogue the
insect diversity of an area. Herein, we provide checklist of forensically
important rove beetles from India belonging to subfamily Staphylininae of
family Staphylinidae.
Keywords: Beetle diversity,
carcass, forensic entomology, Staphylininae, Rove beetles.
Introduction
Diptera and
Coleoptera are the major insect orders which help forensic entomologists in
solving homicidal cases (Catts & Haskell 1990). Generally, dipteran flies
are the first to arrive and colonize a carcass thereby acting as key players in
calculating the post-mortem interval of a body (Bharti & Singh 2003).
Beetles, on the other hand, are attracted to a carcass during the advanced
stage of its decomposition. Silphidae (Carrion beetles or burying beetles) lay
eggs in the burrows created under the carrion and uses carrion as a resource to
complete its life history, Staphylinidae (Rove beetles), Histeridae (Clown
beetles) are predacious and prefer to feed on fly larvae and dead flies present
on the carcass, whereas Dermestid beetles prefer skin of the carcass, and
Trogidae feed on dried remains (Weidner et al. 2021).
Of all the
forensically important families of beetles, Staphylinidae is the most diverse
family of rove beetles with 35 subfamilies encompassing 4,038 genera and 66,928
species (https://www.catalogueoflife.org/). Subfamily Staphylininae with 9,071
species is the third largest subfamily and comprises of nine tribes
(Arrowinini, Coomaniini, Diochini, Maorothini, Othini, Platyprosopini,
Staphyliini, Thayeralinini (extinct), and Xantholiini) of which Staphyliini constitutes
the most hyperdiverse group with 6,377 species (Brunke et al. 2019). Its
members prefer to feed on fungi, dead organic matter, carrion making them a
potential candidate in solving PMI cases. A lot has been done to catalogue the
forensically important beetles (Pajni & Kumar 1977; Kulshrestha &
Satpathy 2001; Bharti & Singh 2003; Dekeirsschieter et al. 2013; Sharma et
al. 2018; Al-Zahrani et al. 2023; Benny & Suleiman 2023; Luo & Meng
2023) but there is a dearth of comprehensive checklist of forensically
important rove beetles from India. Cameron (1943, 1944) initiated the work on
Staphylinidae from India, followed by inputs from Scheerpeltz 1933; Coiffait
1982, 1984; Schillhammer 2001, 2003. But the change in taxonomic status of many
taxa, additions of new records and species in recent times (Smetana 1975, 1988;
Herman 2001; Assing 2008; Zyla & Solodovnikov 2019; Tihelka et al. 2020),
has left the data on rove beetles fragmented and confusing. Thus, in the light
of above stated facts, an updated checklist of forensically important rove
beetles from India belonging to subfamily Staphylininae of family Staphylinidae
is provided.
Results and Discussion
A total of nine
tribes of Staphylininae (including extinct tribe Thayeralinini) are known from
the World. Cameron (1943) divided Indian Staphylininae into three tribes namely
Staphylinini, Xantholinini, and Quediini but phylogenetic analysis revealed
Quediini as polyphyletic assemblage of species belonging to different subtribes
within Staphylinini (Solo-dovnikov 2006; Chatzimanolis et al. 2010; Salnitska
& Solodovnikov 2018; Brunke et al. 2019).
Currently, Staphylininae is represented by tribes Othiini, Staphylinini,
and Xantholinini from India. Tribe Othiini is represented by five genera, 152
species; tribe Staphylinini by 247 genera, 6377 species and tribe Xantholini by
146 genera and 2352 species from the world (https://www.catalogueoflife.org/).
From India, tribe Othiini is represented by genus Othius; Staphylinini
by Philonthus, Quedius, Platydracus, Belonuchus, Bisnius,
Heterothops, Staphylinus, Creophilus, Ontholestes
and Neobisnius where as Xantholinini by Indolinus and Indomorphus.
Following is the list of forensically significant rove beetles of subfamily
Staphylininae from India.
Subfamily:
Staphylininae Latreille, 1802
1802. Staphylininae
Latreille, Familles naturelles des genres. Vol. 3. F. Dufart, Paris. xii
+ 13-467 pp.
Tribe: Othiini
Thomson, C.G., 1859
1859. Othiini
Thomson, C.G., Berlingska Boktryckeriet, Lund 290 pp.
1957. Atrecini Hatch,
University of Washington Publications in Biology 16(2).
Genus: Othius Stephens,
1829
1829. Othius Stephens,
Baldwin and Cradock, London, 68 pp.
1906. Othiellus Casey,
Transactions of the Academy of Science of St. Louis.
1926. Othius
apicalis Cameron, Transactions of the Entomological Society of
London, 1925: 341–372.
Distribution: India
(Sikkim: Darjeeling, Uttar Pradesh), Nepal.
Othius cachemiricus Coiffait, 1982
1982. Othius
cachemiricus Coiffait, Entomologica Basiliensia, 7: 231–302.
Distribution: India
(Kashmir).
Othius flavocaudatus Cameron, 1926
1926. Othius
flavocaudatus Cameron, Transactions of the Entomological Society of
London, 1925: 341–372.
Distribution: India,
Nepal.
Othius kashmiricus Cameron, 1943
1943. Othius kashmiricus
Cameron, Proceedings of the Royal Entomological Society of London (B),
12(1): 127–132.
Distribution: India
(Kashmir).
Othius monticola Cameron, 1943
1943. Othius
monticola Cameron, Proceedings of the Royal Entomological Society of
London (B), 12(1): 127–132.
Distribution: India
(Darjeeling).
Othius pokharensis Coiffait, 1983
1983. Othius
pokharensis Coiffait, Nouvelle Revue d’Entomologie, 13(2):
161–172.
Distribution: India
(Sikkim: Darjeeling), Nepal.
Othius ruficornis Cameron, 1928
1928. Othius
ruficornis Cameron, Annals and Magazine of Natural History, 2(12) :
558–569.
Distribution: India
(Sikkim).
Othius sinuosus Assing, 1998
1998. Othius
sinuosus Assing, Beiträge zur Entomologie, 48(2): 343–351.
Distribution: India, Pakistan (Karakorum).
Othius uttaricus Assing, 2008
2008. Othius
uttaricus Assing, Koleopterologische Rundschau, 78: 245–263.
Distribution: India (Uttarakhand).
Othius yusmargensis Coiffait, 1982
1982. Othius
yusmargensis Coiffait, Entomologica Basiliensia, 7: 231–302.
Distribution: India
(Kashmir).
Tribe: Staphylinini Latreille, 1802
1802. Staphylinini
Latreille, Familles naturelles des genres. Vol. 3. F. Dufart, Paris. xii
+ 13–467pp.
Genus: Belonuchus
Nordmann, 1837
1837. Belonuchus Nordmann,
Academiae Caesareae Scientiarum, Petropoli [St.
Petersburg], 167 pp.
Belonuchus aeneipennis Fauvel, 1895
1895. Belonuchus
aeneipennis Fauvel, Revue d’Entomologie, 14(6–7): 180–286.
Distribution:
Throughout India, Myanmar, Malaysia (Melaka, Sarawak), Singapore, Indonesia
(Borneo, Sumatra, Moluccas).
Belonuchus punctifrons (Cameron, 1926)
1926. Philonthus
punctifrons Cameron, Transactions of the Entomological Society of London,
1925: 341–372.
Distribution: India
(Himachal Pradesh, Uttar Pradesh), Thailand, China (Shaanxi, Sichuan, Yunnan),
Malaysia (Pahang), Indonesia (Sumatra).
Genus: Bisnius Stephens, 1829
1829. Bisnius Stephens,
Baldwin and Cradock, London, 68 pp.
1859. Gefyrobius Thomson,
C.G., Berlingska Boktryckeriet, Lund, 290 pp.
Bisnius deuvei (Coiffait,
1982)
1982. Bisnius
deuvei (Coiffait), Entomologica Basiliensia, 7: 231–302.
Distribution: India
(Kashmir)
Bisnius diversus (Schubert, K.,
1906)
1906. Bisnius
diversus (Schubert, K.,), Deutsche Entomologische Zeitschrift,
(2): 378–383.
1906. Philonthus
diversus Schubert, K., Deutsche Entomologische Zeitschrift, (2):
378–383.
Distribution: India
(Kashmir), Mongolia.
Bisnius fimetarius (Gravenhorst,
1802)
1802. Bisnius
fimetarius (Gravenhorst), Brunsuigae [Braunschweig]. lxvi + 206 pp
1802. Staphylinus
fimetarius Gravenhorst, Brunsuigae [Braunschweig]. lxvi + 206 pp.
1802. Staphylinus
rigidicornis Gravenhorst, Brunsuigae [Braunschweig]. lxvi + 206 pp.
1858. Gabrius
longulus Motschulsky, Bulletin de la Société Impériale des Naturalistes
de Moscou, 31(2): 634–670.
1859. Philonthus
xanthomerus Kraatz, Archiv für Naturgeschichte, 25(1): 1–96.
1860. Philonthus
interpunctatus Hochhuth, Bulletin de la Société Impériale des
Naturalistes de Moscou, 33(2): 539–588.
1861. Philonthus
stenoderus Reiche, Annales de la Société Entomologique de France,
(4)1: 201–210.
Distribution: India
(Arunachal Pradesh, Uttar Pradesh, Uttarakhand), Europe, Russia (European,
Siberia, Far East), Armenia, Georgia, Morocco, Algeria, Tunisia, Libya, Egypt,
Turkey, Iran, Kazakhstan China (Ningxia), Taiwan, Canada.
Bisnius nichinaiensis (Coiffait,
1982)
1982. Bisnius
nichinaiensis (Coiffait), Senckenbergiana Biologica, 62: 21–179.
1982. Philonthus
nichinaiensis Coiffait, Senckenbergiana Biologica, 62: 21–179.
Distribution: India
(Kashmir), China (Yunnan).
Bisnius protenus (Schubert, K.,
1906)
1906. Bisnius
protenus (Schubert, K.,) Deutsche Entomologische Zeitschrift, (2):
378–383.
1906. Philonthus
protenus Schubert, K., Deutsche Entomologische Zeitschrift, (2):
378–383.
1908. Philonthus
signifrons Schubert, K, Deutsche Entomologische Zeitschrift, (5):
609–625.
1911. Philonthus
simlaensis Bernhauer, Entomologische Blätter, 7(2–3): 199–200.
1928. Philonthus
yatungensis Cameron, Annals and Magazine of Natural History, 2(12): 558–569.
Distribution:
Pakistan, India (Kashmir, Himachal Pradesh, Uttar Pradesh), Nepal, China
(Tibet).
Bisnius schawalleri (Coiffait,
1982)
1982. Bisnius
schawalleri (Coiffait), Senckenbergiana Biologica, 62: 21–179.
1982. Philonthus
schawalleri Coiffait, Senckenbergiana Biologica, 62: 21–179.
Distribution: India
(Kashmir).
Genus: Creophilus Leach, 1819
1819. Creophilus Leach,
Thomas Boys, London, 496 pp.
1839.
Creophagus Streubel, Isis von Oken, (2).
1839.
Saprophilus Streubel, Isis von Oken, (2).
Creophilus flavipennis (Hope, 1831)
1831. Creophilus
flavipennis (Hope), The Zoological Miscellany, 1: 21–32.
1859. Creophilus
villipennis Kraatz, Archiv für Naturgeschichte, 25(1): 1–96.
1878. Emus
violaceus Fauvel, Annali del Museo Civico di Storia Naturale di Genova,
12: 171–315.
1879. Emus
insularis Fauvel, Annali del Museo Civico di Storia Naturale di Genova,
15: 63–121.
Distribution:
Northern India, Ryukyu Islands, Nepal, Sri Lanka, Myanmar, Thailand, China
(Qinghai, Yunnan), Taiwan, Laos, Vietnam, Malaysia (Sarawak), Philippines,
Indonesia (Sumatra, Kalimantan, Lombok, Ceram, Buru, Moluccas, Nais, Tanimbar);
New Guinea.
Creophilus maxillosus (Linnaeus, 1758)
1758. Creophilus
maxillosus (Linnaeus), Editio decima, reformata. Tomus I. L. Salvii,
Holmiae. 824 pp.
1758. Staphylinus
maxillosus Linnaeus, Editio decima, reformata. Tomus I. L. Salvii,
Holmiae. 824 pp.
Distribution:
Throughout India, Canada, USA, Mexico, Guatemala, Honduras, Galapagos Is.,
Cuba, Isla de Pinos, Jamaica, Iran, Pakistan, Azores, Canary Islands, Madeira,
Iceland, Greenland, St. Helena, Bhutan, Taiwan, Ryukyu Islands, China, Hawaiian
Islands, Brazil, Chile, Argentina, Peru.
Creophilus maxillosus maxillosus
(Linnaeus, 1758)
1874. Creophilus
imbecillus Sharp, Transactions of the Entomological Society of London,
(1): 1–103.
1899. Creophilus
pulchellus Meier, Entomologische Nachrichten, 25: 97–102.
1908. Creophilus canariensis Bernhauer, Münchener
Koleopterologische Zeitschrift, 3(3): 320–335.
1927. Creophilus sikkimensis Wendeler, Neue
Beiträge zur Systematischen Insektenkunde, 4(1): 1–2.
Distribution: India (Sikkim), Açôres, Canary Islands,
Madeira, Iceland, China, Greenland, Brazil, Chile, Argentina, Bolivia, Peru.
Genus: Heterothops Stephens, 1829
1829. Heterothops Stephens,
Baldwin and Cradock, London, 68 pp.
1837.Trichopygus Nordmann,
Academiae Caesareae Scientiarum, Petropoli [St. Petersburg], 167 pp.
Heterothops dissimilis (Gravenhorst, 1802)
1802. Heterothops dissimilis (Gravenhorst), C.
Reichard, Brunsuigae [Braunschweig], lxvi + 206 pp.
1876. Heterothops
paradoxus Mulsant & Rey, Annales de la Société d’Agriculture
Histoire Naturelle et Arts Utiles de Lyon, 4(8J 145–856.
1977. Heterothops
fraudulenta Coiffait, Nouvelle Revue d’Entomologie, 7(2): 133–143.
1977. Heterothops
armeniaca Coiffait, Nouvelle Revue d’Entomologie, 7(2):
133–143.
1802. Tachyporus
dissimilis Gravenhorst, C. Reichard, Brunsuigae [Braunschweig], 206
pp.
Distribution: India
(Kashmir), Greece, Canary Islands, Morocco, Algeria, Tunisia, Libya, Armenia,
Azerbaijan, Turkey, Iran, Kazakhstan, Europe.
Heterothops franzi Coiffait, 1982
1982. Heterothops
franzi Coiffait, Senckenbergiana Biologica, 62: 21–179.
Distribution: India, Nepal.
Heterothops hindustanus Cameron, 1932
1932. Heterothops
hindustanus Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India
(Tamil Nadu: Nilgiri hills).
Heterothops indicus Cameron, 1926
1926. Heterothops
indicus Cameron, Transactions of the Entomological Society of London,
1925: 341–372.
Distribution:
Throughout India.
Heterothops khairo Smetana, 1988
1988. Heterothops
khairo Smetana, Quaestiones Entomologicae, 24(2): 163–464.
Distribution: India
(Uttar Pradesh).
Heterothops oculatus Fauvel, 1895
1895. Heterothops
oculatus Fauvel, Revue d’Entomologie, 14(6–7): 92–127.
Distribution: India,
Pakistan, Nepal, Myanmar, Taiwan.
Heterothops persimilis Cameron, 1932
1932. Heterothops
persimilis Cameron, Taylor & Francis, London, xiii + 443 pp.
Distribution: India,
Nepal, Myanmar.
Heterothops pusillus Coiffait, 1982
1982. Heterothops
pusillus Coiffait, Senckenbergiana Biologica, 62: 21–179.
Distribution: India
(Himachal Pradesh, Uttar Pradesh), Nepal.
Heterothops saano Smetana, 1988
1988. Heterothops
saano Smetana, Quaestiones Entomologicae, 24(2): 163–464.
Distribution: India
(Kashmir).
Genus: Neobisnius Ganglbauer, 1895
1895. Neobisnius Ganglbauer,
Carl Gerold’s Sohn, Vienna. vi + 881 pp.
Neobisnius cameroni Wendeler, 1928
1928. Neobisnius
cameroni Wendeler, Deutsche Entomologische Zeitschrift, 1928: 37.
1926. Neobisnius fraternus
Cameron, Transactions of the Entomological Society of London, 1925:
341–372.
Distribution: India.
Neobisnius praelongus (Gemminger
& Harold, 1868)
1868. Neobisnius
praelongus (Gemminger & Harold), E.H. Gummi, Monachii [Munich].
pp. 425–752.
1859. Philonthus
longulus Kraatz, Archiv für Naturgeschichte, 25(1): 1–96.
Distribution: India
(Bihar, Nagaland, Sikkim, Uttar Pradesh, West Bengal, Andaman Island), Sri
Lanka, Myanmar, Thailand, Cambodia, Vietnam, Malaysia, Indonesia, Philippines,
Taiwan, China (Zhejiang).
Genus: Ontholestes Ganglbauer, 1895
1833. Trichoderma Stephens,
Baldwin and Cradock, London. iv + 134 pp.
1895.
Ontholestes Ganglbauer, Carl Gerold’s Sohn, Vienna.
vi + 881 pp.
Ontholestes aurosparsus (Fauvel, 1895)
1895. Ontholestes aurosparsus
(Fauvel), Revue d’Entomologie, 14(6–7): 92–127.
1895. Leistotrophus
aurosparsus Fauvel, Revue d’Entomologie, 14(6–7): 92–127.
1936. Ontholestes
ophthalmicus Kirshenblat, Trudy Zoologicheskogo Instituta Akademii Nauk
SSSR, 3: 551–566.
1944. Ontholestes
assamensis Cameron, Proceedings of the Royal Entomological Society of
London (B), 13(1–2): 104–108.
Distribution: India
(Kashmir, Assam, Uttar Pradesh), Nepal, Myanmar, China (Fujian, Guangxi,
Shandong).
Ontholestes tenuicornis (Kraatz, 1859)
1859. Ontholestes
tenuicornis (Kraatz), Archiv für Naturgeschichte, 25(1): 1–96.
1859. Leistotrophus
tenuicornis Kraatz, Archiv für Naturgeschichte, 25(1): 1–96.
Distribution: India
(West Bengal: Chakrata District), Nepal, China (Fujian).
Genus: Philonthus Stephens, 1829
1829. Philonthus Stephens,
Baldwin and Cradock, London, 68 pp.
1968. Endeius Coiffait
& Sáiz, Centre National de la Recherche Scientifique, Paris,
472 pp.
1977. Philonthus
(Parambigus) Marcuzzi, Quaderni di Ecologia Animale, 12.
Philonthus (Onychophilonthus)
anepsius Cameron, 1926
1926. Philonthus
anepsius Cameron, Transactions of the Entomological Society of London,1925:
341–372.
Distribution: India
(Himachal Pradesh).
Philonthus (Onychophilonthus)
funeralis Cameron, 1943
1943. Philonthus
funeralis Cameron, Proceedings of the Royal Entomological Society of
London (B), 12(1): 127–132.
Distribution: India
(Sikkim: Darjeeling).
Philonthus (Onychophilonthus)
kashmiranus Bernhauer, 1915
1915. Philonthus
kashmiranus Bernhauer, Coleopterologische Rundschau, 4(3): 49–60.
Distribution: India
(Kashmir), Nepal.
Philonthus (Philonthus)
aberrans Cameron, 1932
1932. Philonthus
aberrans Cameron, Fauna of British India, including Ceylon and Burma.
Taylor & Francis, London. xiii + 443 pp.
Distribution: India
(Uttar Pradesh).
Philonthus (Philonthus)
acroleucus Kraatz, 1859
1859. Philonthus
acroleucus Kraatz, Archiv für Naturgeschichte, 25(1): 1–96.
Distribution: India,
Sri Lanka, Philippines, Indonesia (Sumatra), China (Hong Kong).
Philonthus (Philonthus)
adversus Bernhauer & K.Schubert, 1914
1914. Philonthus
adversus Bernhauer & K. Schubert, Coleopterorum Catalogus, Pars 57. W.
Junk, Berlin, 289–408 pp.
1925.Philonthus
adversus Cameron, Catalogue of the Indian Insects, 126 pp.
Distribution: India
(Sikkim), Myanmar.
Philonthus (Philonthus)
aeger Eppelsheim, 1895
1895.Philonthus
aeger Eppelsheim, Wiener Entomologische Zeitung, 14(3): 53–70.
Distribution: India
(Himachal Pradesh, Manipur, Uttar Pradesh), Nepal, Bhutan, China (Sichuan),
Vietnam.
Philonthus (Philonthus)
aeneipennis Boheman, 1858
1858.Philonthus
aeneipennis Boheman, Insecta, 1–112.
1908.Philonthus kuluensis Schubert, K., Deutsche
Entomologische Zeitschrift, 1908 (5): 609–625.
1983. Philonthus
arachosicus Coiffait, Bonner Zoologische Beiträge, 34: 477–483.
Distribution: India,
Saudi Arabia, Iran, Afghanistan, Pakistan, Nepal, Bhutan, Andaman Is., Vietnam,
China, Taiwan, Ryukyu Is., Philippines, Indonesia, Japan (Honshu, Kyushu,
Shikoku), Korea (S), Ogasawara Is., New Guinea, Yemen, Tanzania, Senegal,
Mauritius.
Philonthus (Philonthus)
aeripennis Cameron, 1943
1943. Philonthus
aeripennis Cameron, Proceedings of the Royal Entomological Society of
London (B), 12(1): 127–132.
Distribution: India
(Darjeeling).
Philonthus (Philonthus)
aliquatenus Schubert, K., 1908
1908. Philonthus aliquatenus Schubert, K., Deutsche
Entomologische Zeitschrift, 1908 (5): 609–625.
Distribution: India
(Kashmir, Himachal Pradesh, Manipur, Uttar Pradesh), Nepal, Bhutan, China
(Yunnan).
Philonthus (Philonthus)
amplitarsis Schillhammer, 2001
2001. Philonthus
amplitarsis Schillhammer, Koleopterologische Rundschau, 71: 59–61.
Distribution: India
(Uttar Pradesh).
Philonthus (Philonthus)
assamensis Cameron, 1932
1932.Philonthus
assamensis Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India
(Assam, Manipur, Meghalaya, Sikkim, Tripura, Uttarakhand).
Philonthus (Philonthus)
atricoxis Cameron, 1943
1943.Philonthus
atricoxis Cameron, Proceedings of the Royal Entomological Society of
London (B), 12(1): 127–132.
Distribution: India
(Kashmir).
Philonthus (Philonthus)
azuripennis Cameron, 1928
1928.Philonthus
azuripennis Cameron, Annals and Magazine of Natural History, (10)
2(12): 558–569.
1982.Philonthus
trisulensis Coiffait, Senckenbergiana Biologica, 62: 21–179.
Distribution: India
(Sikkim), Nepal, Bhutan, China (Gansu, Qinghai, Shaanxi, Sichuan, Yunnan,
Xizang).
Philonthus (Philonthus)
batotensis Cameron, 1932
1932.Philonthus
batotensis Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India
(Kashmir), Afghanistan.
Philonthus (Philonthus)
beesoni Cameron, 1926
1926.Philonthus
beesoni Cameron, Transactions of the Entomological Society of London,
1925: 341–372.
Distribution: India
(Himachal Pradesh, Uttar Pradesh), Afghanistan, Pakistan, Nepal.
Philonthus (Philonthus)
bengalensis Bernhauer, 1911
1911. Philonthus
bengalensis Bernhauer, Entomologische Blätter, 7(2–3):
199–200.
Distribution: India
(Bihar, Tamil Nadu, Uttar Pradesh).
Philonthus (Philonthus)
birmanus Fauvel, 1895
1895. Philonthus
birmanus Fauvel, Revue d’Entomologie, 14(6–7): 92–127.
Distribution: India,
Myanmar, China (Xizang), Vietnam, Indonesia (Sumatra).
Philonthus (Philonthus)
brevithorax Bernhauer, 1934
1934. Philonthus
brevithorax Bernhauer, Entomologisches Nachrichtenblatt (Troppau),
8(1): 1–20.
Distribution: India
(Himachal Pradesh, Uttar Pradesh), Nepal, Bhutan, China (Sichuan).
Philonthus (Philonthus)
carbonarius (Gravenhorst, 1802)
1802. Philonthus (Philonthus)
carbonarius (Gravenhorst), C. Reichard, Brunsuigae [Braunschweig].
lxvi + 206 pp.
1832. Philonthus
nigroaeneus Stephens, Mandibulata, Vol. 5. Baldwin & Cradock,
London, 240 pp.
1857. Philonthus
nemorosus Gistel, Schorner’schen Buchhandlung, Straubing, 94 pp.
1905.Philonthus
shetlandicus Poppius, Öfversigt af Finska Vetenskaps-Societetens
Förhandlingar, 47(17): 1–19.
1933. Philonthus
menetriesi Kirshenblat, Entomologicheskoe Obozrenie, 25(1–2).
1952. Philonthus
vinohradensis Dvořák, R. & Havelka, Bulletin de la Société
Entomologique de Mulhouse, 41.
Distribution: India,
Europe, Russia (European, eastern & western Siberia, Far East), Algeria,
Canary Islands, Armenia, Georgia, Turkey, Iran, Kazakhstan, Kyrgyzstan,
Uzbekistan, Nepal, Mongolia, China (Heilongjiang), Canada, USA.
Philonthus (Philonthus)
caucasicus Nordmann, 1837
1837. Philonthus
caucasicus Nordmann, Academiae Caesareae Scientiarum, Petropoli [St.
Petersburg]. 167 pp.
1851. Philonthus
rutilipennis Hochhuth, Bulletin de la Société Impériale des Naturalistes
de Moscou, 24(2): 3–58.
1900. Philonthus
apicalis Leinberg, Meddelanden af Societas pro Fauna et Flora Fennica,
26: 79–80.
1913. Philonthus
heinemanni Bernhauer, Coleopterologische Rundschau, 2(8–9): 130–134.
Distribution: India
(Kashmir), Europe, Russia (European, eastern & western Siberia), Armenia,
Georgia, Turkey, Syria, Iraq, Iran, Afghanistan, Kazakhstan, Tajikistan,
Turkmenistan, Uzbekistan, Mongolia, Canada, USA.
Philonthus (Philonthus)
chatterjeei Cameron, 1926
1926.Philonthus
chatterjeei Cameron, Transactions of the Entomological Society of London,
1925: 341–372.
Distribution: India
(Uttar Pradesh, Uttarakhand).
Philonthus (Philonthus)
christiei Bernhauer, 1920
1920. Philonthus christiei Bernhauer, Archiv
für Naturgeschichte, 84(A)(10): 177–184.
1926. Philonthus
masuriensis Cameron, Transactions of the Entomological Society of London,
1925: 341–372.
Distribution: India
(Himachal Pradesh, Uttar Pradesh).
Philonthus (Philonthus)
cinctipennis Fauvel, 1875
1875.Philonthus
cinctipennis Fauvel, F. le Blanc-Hardel, Caen, 3: 1–38.
Distribution: India,
Egypt, Oman, Iraq, Ethiopia, Somalia, Myanmar, Vietnam, Indonesia (Sumatra),
Philippines.
Philonthus (Philonthus)
cinctulus (Gravenhorst, 1802)
1802. Philonthus (Philonthus)
cinctulus (Gravenhorst), Brunsuigae [Braunschweig]. lxvi + 206 pp.
Distribution: India
(Uttar Pradesh, Uttarakhand, West Bengal), Pakistan, Nepal, Sri Lanka, Myanmar,
Vietnam.
Philonthus (Philonthus)
cliens Eppelsheim, 1890
1830. Philonthus
cliens Eppelsheim, Wiener Entomologische Zeitung, 9(10): 217–229.
1960. Philonthus
lindemanni Scheerpeltz, Opuscula Zoologica (München), 51: 1–7.
Distribution: India,
Pakistan, Nepal, Botswana, Senegal, Tanzania.
Philonthus (Philonthus)
cognatus Stephens, 1832
1832. Philonthus
cognatus Stephens, Mandibulata, Vol. 5. Baldwin & Cradock, London,
240pp.
1832. Philonthus
melanopterus Stephens, Mandibulata, Vol. 5. Baldwin & Cradock,
London, 240 pp.
1832. Philonthus
nigripennis Stephens, Mandibulata, Vol. 5. Baldwin
& Cradock, London, 240 pp.
Distribution: India
(Kashmir), Algeria, Morocco, Russia (European, eastern & western Siberia),
Tunisia, Madeira, Turkey, Armenia, Iran, Kazakhstan, Kyrgyzstan, Tajikistan,
Pakistan, Nepal, China (Jilin), Sri Lanka, Canada, USA.
Philonthus (Philonthus) concinnus (Gravenhorst, 1802)
1802. Philonthus (Philonthus)
concinnus (Gravenhorst), Brunsuigae [Braunschweig]. lxvi + 206 pp.
1830. Staphylinus
irregularis Mannerheim, Précis d’un nouvel arrangement de la famille des
brachélytres, de l’ordre des insectes coléoptères. St. Petersbourg, 87 pp.
1876. Philonthus
melanarius Mulsant & Rey, Annales de la Société d’Agriculture
Histoire Naturelle et Arts Utiles de Lyon, 4(8): 145–856.
1895. Philonthus
minor Ganglbauer, Carl Gerold’s Sohn, Vienna. vi + 881 pp.
1910. Philonthus
ochripennis Gerhardt, Deutsche Entomologische Zeitschrift, (5):
554–557.
Distribution: India
(Sikkim), Europe, Algeria, Morocco, Libya, Azores, Canary Islands, Russia (Eastern & western Siberia, Far East),
Armenia, Georgia, Turkey, Lebanon, Israel, Iran, Afghanistan, Kazakhstan,
Kyrgyzstan, Tajikistan, Uzbekistan, Mongolia, Canada, USA.
Philonthus (Philonthus)
congruens Cameron, 1932
1932. Philonthus
congruens Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India
(Uttar Pradesh).
Philonthus (Philonthus)
convalescens Eppelsheim, 1890
1890. Philonthus
convalescens Eppelsheim, Wiener Entomologische Zeitung, 9(10):
161–172.
Distribution: India
(Himachal Pradesh, Sikkim, Uttar Pradesh), Nepal, China (Xizang).
Philonthus (Philonthus)
crassicornis Fauvel, 1895
1895. Philonthus
crassicornis Fauvel, Revue d’Entomologie, 14(6–7): 92–127.
Distribution: India
(Tamil Nadu, Uttar Pradesh, Uttarakhand), Sri Lanka, Myanmar, Vietnam,
Singapore, Indonesia (Java, Sumatra), Taiwan, Madagascar, Mauritius,
Philippines.
Philonthus (Philonthus)
cruentatus (Gmelin, 1790)
1790. Philonthus (Philonthus)
cruentatus (Gmelin), G. E. Beer, Lipsiae, 1(4): 1517–2224.
1900. Philonthus
extinctus Bernhauer, Wiener Entomologische Zeitung, 19(2–3): 46–55.
1924. Philonthus
immaculatus Gusmann, Entomologische Blätter, 20(3): 248–252.
Distribution: India
(Uttar Pradesh), Europe, Russia (Eastern & western Siberia, Far East),
Georgia, Armenia, Algeria, Morocco, Canary Islands, Turkey, Lebanon, Iran,
Kazakhstan, Kyrgyzstan, Afghanistan, Tajikistan, Uzbekistan, Mongolia, Japan,
Canada, USA, Mexico.
Philonthus (Philonthus)
cyanelytrius Kraatz, 1859
1859. Philonthus cyanelytrius Kraatz, Archiv
für Naturgeschichte, 25(1): 1–96.
1926. Hesperus gridellii Cameron, Transactions
of the Entomological Society of London, 1925: 341–372.
Distribution: India (Kashmir, Himachal Pradesh,
Meghalaya, Uttar Pradesh), Nepal, Vietnam.
Philonthus (Philonthus)
dejectus Cameron, 1932
1932. Philonthus
dejectus Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India
(Uttar Pradesh), Nepal.
Philonthus (Philonthus)
delicatulus Boheman, 1858
1858. Philonthus
delicatulus Boheman, Insecta, 1–112.
1858. Philonthus
lativentris Motschulsky, Bulletin de la Société Impériale des
Naturalistes de Moscou, 31 (2): 204–264.
1859. Philonthus
subirideus Kraatz, Archiv für Naturgeschichte, 25(1): 1–96.
Distribution: India
(Uttarakhand), China (Hong Kong), Sri Lanka, Myanmar, Vietnam, Singapore, Indonesia
(Java, Sumatra); Philippines, Taiwan.
Philonthus (Philonthus)
densus Cameron, 1926
1926. Philonthus
densus Cameron, Transactions of the Entomological Society of London,
1925: 341–372.
Distribution: India
(Uttar Pradesh), Afghanistan, Nepal, China (Yunnan).
Philonthus (Philonthus)
distincticornis Cameron, 1932
1932. Philonthus distincticornis Cameron, Taylor
& Francis, London. xiii + 443 pp.
1932. Philonthopsis antennalis Cameron, Taylor
& Francis, London. xiii + 443 pp.
1991. Philonthus cyaneonitens Schillhammer, Koleopterologische
Rundschau, 61: 51–56.
Distribution: India (Sikkim, Uttar Pradesh), Nepal.
Philonthus (Philonthus) donckieri
Bernhauer, 1915
1915. Philonthus donckieri Bernhauer,
Entomologische Blätter, 11(10–12): 251–258.
1919. Philonthus tripunctatus Cameron,
Entomologische Blätter, 11(10–12): 251–258.
1931. Philonthus crebrior Bernhauer,
Entomologische Zeitung, 48(3): 125–132.
Distribution: India (Goa, Himachal Pradesh, Tamil
Nadu, Uttar Pradesh), Nepal, Sri Lanka, Laos, Vietnam, China (Fujian,
Guangdong, Hong Kong, Jiangxi, Sichuan).
Philonthus (Philonthus) explorator
Cameron, 1932
1932. Philonthus explorator Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India (Tamil Nadu: Nilgiri Hills),
Nepal, Thailand, Taiwan, China (Shanghai).
Philonthus (Philonthus)
flavipes Kraatz, 1859
1859. Philonthus flavipes Kraatz, Archiv für
Naturgeschichte, 25(1): 1–96.
Distribution: India
(Sikkim: Darjeeling, Uttarakhand, Manipur, Tamil Nadu), China (Fujian, Hong
Kong, Yunnan, Zhejiang), Taiwan, Ryukyu Islands, Sri Lanka, Myanmar, Thailand,
Vietnam, Indonesia (Sumatra), Brunei, Samoa, Réunion, Mauritius, Philippines.
Philonthus (Philonthus)
flavocinctus Motschulsky, 1858
1858. Philonthus flavocinctus Motschulsky, Bulletin
de la Société Impériale des Naturalistes de Moscou, 31(2): 204–264.
1859. Philonthus rufo-marginatus Kraatz, Archiv
für Naturgeschichte, 25(1): 1–96.
Distribution: India
(West Bengal), Israel, China (Hong Kong), Taiwan, Ryukyu Islands, Ogasawara
Islands, Sri Lanka, Myanmar, Vietnam, Singapore, Indonesia (Sumatra), New
Caledonia, Sudan, Réunion, Mauritius, Réunion, Rodriguez; Philippines, Senegal,
Madagascar.
Philonthus (Philonthus)
fletcheri Cameron, 1943
1943. Philonthus
fletcheri Cameron, Proceedings of the Royal Entomological Society of
London (B), 12(1): 127–132.
Distribution: India
(Kashmir).
Philonthus (Philonthus)
foetidus Cameron, 1932
1932. Philonthus (Philonthus)
foetidus Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India (Himachal Pradesh, Uttar Pradesh),
Nepal, Myanmar, Thailand, Taiwan, China (Shanghai).
Philonthus (Philonthus)
fuscatus Kraatz, 1859
1859. Philonthus fuscatus Kraatz, Archiv für
Naturgeschichte, 25(1): 1–96.
Distribution: India, Iraq, Sri Lanka, China.
Philonthus (Philonthus)
gardneri Cameron, 1932
1932. Philonthus gardneri Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India (Sikkim, Uttarakhand).
Philonthus (Philonthus)
gemellus Kraatz, 1859
1859. Philonthus gemellus Kraatz, Archiv für
Naturgeschichte, 25(1): 1–96.
Distribution: India (Uttar Pradesh, West Bengal),
Pakistan, China (Beijing, Hong Kong), Ryukyu Islands (Ishigaki), Sri Lanka,
Singapore, Indonesia (Java), Réunion, Mauritius, Madagascar.
Philonthus (Philonthus)
geminus Kraatz, 1859
1859. Philonthus geminus Kraatz, Archiv für
Naturgeschichte, 25(1): 1–96.
Distribution: Throughout India, China (Hong Kong,
Yunnan), Ryukyu Islands (Amami-Oshima), Sri Lanka, Myanmar, Thailand, Malaysia,
Vietnam, Singapore, Indonesia.
Philonthus (Philonthus)
gentilicius Cameron, 1926
1926. Philonthus gentilicius Cameron, Transactions
of the Entomological Society of London, 1925: 341–372.
Distribution: India (Himachal Pradesh, Uttar Pradesh).
Philonthus (Philonthus)
gulmargensis Coiffait, 1982
1982. Philonthus gulmargensis Coiffait, Senckenbergiana
Biologica, 62: 21–179.
Distribution: India (Kashmir).
Philonthus (Philonthus)
humilis Cameron, 1932
1932. Philonthus humilis Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India (Uttar Pradesh).
Philonthus (Philonthus)
idiocerus Kraatz, 1859
1859. Philonthus idiocerus Kraatz, Archiv
für Naturgeschichte, 25(1): 1–96.
Distribution: India (Uttar Pradesh), Afghanistan, Sri
Lanka, Nepal, Myanmar, Thailand, Vietnam, China (Hong Kong, Macao, Yunnan);
Indonesia, Philippines.
Philonthus (Philonthus)
incultus Cameron, 1932
1932. Philonthus incultus Cameron, Taylor
& Francis, London. xiii + 443pp.
Distribution: India (Uttar Pradesh).
Philonthus (Philonthus)
industanus Fauvel, 1903
1903. Philonthus industanus Fauvel, Revue
d’Entomologie, 22: 149–163.
1911. Philonthus bipunctatus Bernhauer, Entomologische
Blätter, 7(2–3): 86–93.
1919. Philonthus pubipennis Cameron, Entomologist’s
Monthly Magazine, 55: 251–255.
Distribution: India (Punjab, Sikkim, Tamil Nadu,
Uttarakhand, West Bengal), Nepal, Sri Lanka, Vietnam, China (Hong Kong), w
Malaysia, Indonesia (Sumatra).
Philonthus (Philonthus)
kashmiricus Cameron, 1932
1932. Philonthus kashmiricus Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India (Kashmir), Nepal.
Philonthus (Philonthus)
kempi Cameron, 1924
1924. Philonthus (Philonthus) kempi
Cameron, Records of the Indian Museum, 26(1): 115–122.
Distribution: Pakistan, India (Meghalaya, Uttar
Pradesh), Myanmar.
Philonthus (Philonthus)
kotgarhensis Cameron, 1932
1932. Philonthus (Philonthus)
kotgarhensis Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India (Himachal Pradesh, Uttar Pradesh).
Philonthus (Philonthus)
lidarensis Cameron, 1932
1932. Philonthus (Philonthus)
lidarensis Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India (Kashmir).
Philonthus (Philonthus)
madurensis Bernhauer, 1915
1915. Philonthus madurensis Bernhauer, Entomologische
Blätter, 11(10–12): 251–258.
Distribution: India (Tamil Nadu), Taiwan.
Philonthus (Philonthus)
maindroni Fauvel, 1903
1903. Philonthus maindroni Fauvel, Revue
d’Entomologie, 22: 149–163.
1924. Philonthus annandalei Cameron, Records
of the Indian Museum, 26(1): 118–119.
Distribution: India (Assam, Meghalaya, Sikkim, Tamil
Nadu, Uttar Pradesh, Uttarakhand), Myanmar, Taiwan.
Philonthus (Philonthus)
minutus Boheman, 1848
1848. Philonthus (Philonthus) minutus
Boheman, Officina Norstedtiana, Holmiae. viii + 625 pp.
1874. Philonthus mutans Sharp, Transactions
of the Entomological Society of London, (1): 1–103.
1878. Philonthus rufocinctus Fauvel, Bulletin
de la Société Linnéenne de Normandie, 3(2): 83–162.
1879. Philonthus longiceps Fauvel, Annali
del Museo Civico di Storia Naturale di Genova, 15: 63–121.
Distribution: India, Spain, Morocco, Algeria, Tunisia,
Egypt, Saudi Arabia, Canary Islands, Pakistan, Korea (North & South),
Japan, Senegal, Ethiopia, Kenya, Zaire, Mozambique, South Africa, Sri Lanka,
Myanmar, Vietnam, Malaysia, Indonesia, China, Taiwan, New Guinea, Australia,
Iran.
Philonthus (Philonthus)
montivagans Cameron, 1943
1943. Philonthus montivagans Cameron, Proceedings
of the Royal Entomological Society of London (B), 12(1): 127–132.
Distribution: India (Sikkim: Darjeeling).
Philonthus (Philonthus)
nigricoxis Cameron, 1928
1928. Philonthus (Philonthus) nigricoxis
Cameron, Annals and Magazine of Natural History, 2 (10): 558–569.
Distribution: India (Kashmir, Sikkim), Nepal, China
(Xizang).
Philonthus (Philonthus)
obsoletus Eppelsheim, 1895
1895. Philonthus obsoletus Eppelsheim, Wiener
Entomologische Zeitung, 14(3): 53–70.
1977. Philonthus bhutanicus Coiffait, Entomologica
Basiliensia, 2: 205–242.
1982. Philonthus analokus Coiffait, Senckenbergiana
Biologica, 62: 21–179.
Distribution: India (Himachal Pradesh), Nepal, Bhutan,
Myanmar, China (Xizang).
Philonthus (Philonthus)
paederoides (Motschulsky, 1858)
1858. Philonthus (Philonthus) paederoides
(Motschulsky), Bulletin de la Société Impériale des Naturalistes de Moscou,
31(2): 204–264.
1859. Philonthus bellus Kraatz, Archiv für
Naturgeschichte, 25(1): 1–96.
1922. Philonthus obscuricollis Bernhauer, Archiv
für Naturgeschichte, 88 (A) (10): 147–159.
Distribution: India
(Manipur, Tamil Nadu, West Bengal), Afghanistan, Pakistan, Sri Lanka, Nepal,
Myanmar, Thailand, Cambodia, Vietnam, China (Hong Kong, Yunnan), Taiwan,
Indonesia, Philippines; Malaysia (Sarawak).
Philonthus (Philonthus)
paradoxus Cameron, 1932
1932. Philonthus paradoxus Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India (Uttar Pradesh).
Philonthus (Philonthus)
peregrinus Fauvel, 1866
1866. Philonthus peregrinus Fauvel, Annales
de la Société Entomologique de France, 4(6): 293–340.
Distribution: India (Kashmir, Uttar Pradesh), Nepal,
São Tomé, Senegal, Sierra Leone, Liberia, Ivory Coast, Benin, Cameroon, Central
African Republic, Ethiopia, Uganda, Rwanda, Burundi, Zaire, Kenya, Tanzania,
Namibia, South Africa, Madagascar, Réunion, Mauritius, Seychelles, Comoros.
Philonthus (Philonthus)
persimilis Cameron, 1926
1926. Philonthus persimilis Cameron, Transactions
of the Entomological Society of London, 1925: 341–372.
Distribution: India (Himachal Pradesh, Uttar Pradesh),
Afghanistan.
Philonthus (Philonthus)
politus (Linnaeus, 1758)
1758. Philonthus (Philonthus) politus
(Linnaeus), Tomus I.L. Salvii, Holmiae, 824 pp.
1887. Cafius amblyterus Olliff, Proceedings
of the Linnean Society of New South Wales, (2) 2(3): 471–512.
1923. Quedius ohiaensis Broun, New Zealand
Institute, Bulletin No. 1: 667–708.
1987. Philonthus temperei Coiffait, Nouvelle
Revue d’Entomologie, 3(4): 497–498.
2005. Philonthus rubroelytratus Whitehead, Entomologist’s
Monthly Magazine, 141.
Distribution: India (Karnataka), Europe, Africa,
Canary Islands, Madeira, Azores, Iceland, Russia, Armenia, Azerbaijan, Georgia,
Turkey, Iran, Turkmenistan, Kazakhstan, Mongolia, China, Canada, USA, Mexico,
Venezuela, Chile, Argentina, Australia, New Zealand, Chatham Islands.
Philonthus (Philonthus)
productus Kraatz, 1859
1859. Philonthus (Philonthus)
productus Kraatz, Archiv für Naturgeschichte, 25(1): 1–96.
1931. Philonthus sulciceps Bernhauer, Wiener
Entomologische Zeitung, 48(3): 125–132.
Distribution: India
(Uttar Pradesh), Sri Lanka, Malaysia, China (Hong Kong, southwest), Taiwan,
Ryukyu Islands (Okinawa, Ishigaki, Yonaguni).
Philonthus (Philonthus)
purpuripennis Reitter, 1887
1887. Philonthus (Philonthus)
purpuripennis Reitter, Horae Societatis Entomologicae Rossicae, 21:
201–234.
1933. Philonthus magnificus Bernhauer, Entomologisches
Nachrichtenblatt (Troppau), 7(2): 39–54.
Distribution: India (Andhra Pradesh), Nepal, China
(Gansu, Hubei, Qinghai, Shaanxi, Sichuan, Yunnan, Xinjiang, Xizang).
Philonthus (Philonthus)
remotus Fauvel, 1895
1895. Philonthus (Philonthus) remotus
Fauvel, Revue d’Entomologie, 14(6–7): 264.
1920. Philonthus nilgiriensis Cameron, Entomologist’s
Monthly Magazine, 56: 214–220.
Distribution: India (Himachal Pradesh, Uttar Pradesh),
Myanmar, Indonesia (Java), China (Hong Kong).
Philonthus (Philonthus)
riparius Cameron, 1926
1926. Philonthus (Philonthus)
riparius Cameron, Transactions of the Entomological Society of London,
1925: 341–372.
Distribution: India (Himachal Pradesh, Uttar Pradesh),
Afghanistan, Bhutan, China (Xizang).
Philonthus (Philonthus)
rivularis Cameron, 1932
1932. Philonthus (Philonthus)
rivularis Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India (Kashmir, Himachal Pradesh, Uttar
Pradesh).
Philonthus (Philonthus)
rotundicollis (Ménétriés, 1832)
1832. Philonthus (Philonthus)
rotundicollis (Ménétriés), Académie Impériale des Sciences, St.
Pétersbourg, 271 pp.
1840. Philonthus scutatus Erichson, F.H.Morin,
Berlin, 954 pp.
1860. Philonthus duplopunctatus Motschulsky, Bulletin
de la Société Impériale des Naturalistes de Moscou, 33(2): 539–588.
1916. Philonthus nigropolitus Bernhauer,
Neue Beiträge zur Systematischen Insektenkunde, 1(4): 26–28.
Distribution: India,
Europe, Russia (Siberia, Far East), Turkey, Georgia, Iran, Turkmenistan,
Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, Uzbekistan, Afghanistan,
Pakistan, Mongolia, Japan (Hokkaido), Korea (North & South), China
(Heilongjiang, Qinghai, Shaanxi, Xinjiang).
Philonthus (Philonthus)
rubricollis Motschulsky, 1858
1858. Philonthus (Philonthus)
rubricollis Motschulsky, Bulletin de la Société Impériale des
Naturalistes de Moscou, 31(2): 204–264.
1859. Philonthus erythrostictus Kraatz, Archiv
für Naturgeschichte, 25(1): 1–96.
1935. Philonthus obscuricollis Bernhauer, Entomologisches
Nachrichtenblatt (Troppau), 9(1): 4–14.
Distribution: India (Bihar: Katihar), Myanmar, China
(Fujian, Guangxi, Hong Kong).
Philonthus (Philonthus)
semiaenescens Bernhauer, 1920
1920. Philonthus (Philonthus) semiaenescens
Bernhauer, Archiv für Naturgeschichte, 84 (A)(10): 177–188.
Distribution: India (Uttar Pradesh, Uttarakhand),
Myanmar.
Philonthus (Philonthus)
simpliciventris Bernhauer, 1933
1933. Philonthus (Philonthus)
simpliciventris Bernhauer, Entomologisches Nachrichtenblatt (Troppau),
7(2): 39–54.
1859. Philonthus proximus Kraatz, Archiv für
Naturgeschichte, 25(1): 1–96.
1939. Philonthus explanipes Tottenham, Entomologist’s
Monthly Magazine, 75: 218–224.
Distribution: India (Kashmir, Himachal Pradesh, Uttar
Pradesh), Nepal, China (Beijing, Fujian, Gansu, Shaanxi, Shanxi, Sichuan,
Yunnan), Taiwan, Vietnam.
Philonthus (Philonthus)
siwalikensis Cameron, 1926
1926. Philonthus (Philonthus)
siwalikensis Cameron, Transactions of the Entomological Society of
London, 1925: 341–372.
Distribution: India (Himachal Pradesh, Uttar Pradesh).
Philonthus (Philonthus)
speciosus Cameron, 1926
1926. Philonthus (Philonthus)
speciosus Cameron, Transactions of the Entomological Society of London,
1925: 341–372.
Distribution: India (Uttar Pradesh), Oman, Turkey.
Philonthus (Philonthus)
subjectus Cameron, 1932
1932. Philonthus (Philonthus) subjectus
Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India (Sikkim).
Philonthus (Philonthus)
succicola Thomson, C. G., 1860
1860. Philonthus (Philonthus)
succicola Thomson, C. G., Berlingska Boktryckeriet, Lund, 304 pp.
1835. Staphylinus nigritus Runde, Ploetzianis,
Halae, 32 pp.
1952. Philonthus wohlgrothi Dvořák, R., Bulletin
de la Société Entomologique de Mulhouse, 1952: 41.
Distribution: India (Himachal Pradesh), Europe,
Tunisia, Russia (Siberia, Far East, Kuriles), Armenia, Georgia, Iran,
Kazakhstan, Mongolia, Nepal, China (Heilongjiang, Jilin, Liaoning).
Philonthus (Philonthus)
terminipennis Tottenham, 1939
1939. Philonthus (Philonthus)
terminipennis Tottenham, Proceedings of the Royal Entomological Society
of London (B), 8(12): 227–237.
1932. Philonthus apicipennis Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India (Himachal Pradesh, Uttar Pradesh),
Nepal.
Philonthus (Philonthus)
tricoloris Schubert K., 1908
1908. Philonthus (Philonthus) tricoloris
Schubert, K., Deutsche Entomologische Zeitschrift, (5): 609–625.
Distribution: India (Himachal Pradesh, Uttar Pradesh),
Nepal, Myanmar, Pakistan.
Philonthus (Philonthus)
varians (Paykull, 1789)
1789. Philonthus (Philonthus) varians
(Paykull), Johann. Edman, Upsaliae [Uppsala], 81pp.
Distribution: India,
Europe, Russia, Armenia, Georgia, Turkey, Syria, Iran, Kazakhstan, Kyrgyzstan,
Uzbekistan, Mongolia, Japan, Africa, Afrotropical, South Africa, Madagascar,
Myanmar, Tahiti, Canada, USA, Cuba, St. John, St. Vincent, Venezuela, Argentina,
Chile.
Philonthus (Philonthus)
variipennis Kraatz, 1859
1859. Philonthus (Philonthus)
variipennis Kraatz, Archiv für Naturgeschichte, 25(1): 1–96.
1953. Philonthus
ilus Tottenham, Annals and Magazine of Natural History, 12(6):
143–148.
1979. Philonthus ledouxi
Coiffait, Annales de la Société Entomologique de France, 14: 551–569.
1981. Philonthus
arabicus Coiffait, Fauna of Saudi Arabia, 3: 236–242.
1987. Philonthus
falcozi Coiffait, Nouvelle Revue d’Entomologie, 3(4): 497–498.
Distribution: India
(Uttar Pradesh), Egypt, Saudi Arabia, Afghanistan, Japan (Shikoku), Ryukyu Is.
(Amami-Oshima, Kikai, Kita Daito), Sri Lanka, Thailand, Cambodia, Vietnam,
Malaysia, Indonesia, Philippines, China (Hong Kong), Taiwan.
Philonthus amplicollis Schillhammer, 2003
2003. Philonthus
amplicollis Schillhammer, Koleopterologische Rundschau, 73: 85–136.
Distribution: India (Uttarakhand: Kumaon ), Nepal.
Philonthus andrewesi Cameron, 1920
1920. Philonthus
andrewesi Cameron, Entomologist’s Monthly Magazine, 56: 141–148,
214–220.
Distribution: India (Tamil Nadu).
Philonthus atkinsoni Cameron, 1932
1932. Philonthus
atkinsoni Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India.
Philonthus cameroni Scheerpeltz, 1933
1933. Philonthus cameroni Scheerpeltz, Coleopterorum
Catalogus, Pars 129. W. Junk, Berlin.
1926. Philonthus
fraternus Cameron, Transactions of the Entomological Society of London,
1925: 341–372.
Distribution: India.
Philonthus castaneus Gemminger &
Harold, 1868
1868. Philonthus
castaneus Gemminger & Harold, E. H. Gummi, Monachii [Munich].
pp. 425–752.
1858. Gabrius
badius Motschulsky, Bulletin de la Société Impériale des Naturalistes de
Moscou, 31(2): 204–264.
Distribution: India (Karnataka, Kerala, Manipur,
Uttarakhand, West Bengal).
Philonthus chandigarhensis Pajni &
Kohli, 1977
1977. Philonthus chandigarhensis Pajni &
Kohli, Oriental Insects, 11(4): 513–519.
Distribution: India (Punjab).
Philonthus differens Cameron, 1932
1932.Philonthus differens Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India.
Philonthus dohertyi Cameron, 1932
1932. Philonthus dohertyi Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India (Uttarakhand), Myanmar.
Philonthus flavohumeralis Schillhammer,
2003
2003. Philonthus flavohumeralis Schillhammer, Koleopterologische
Rundschau, 73: 85–136.
Distribution: India, (Sikkim: Darjeeling), Nepal.
Philonthus fulvitarsis Motschulsky, 1860
1860. Philonthus fulvitarsis Motschulsky, Études
Entomologiques, 8: 25–118.
Distribution: India, Sri Lanka.
Philonthus indicus Cameron, 1920
1920. Philonthus indicus Cameron, Entomologist’s
Monthly Magazine, 56: 214–220.
Distribution: India.
Philonthus leptocerus Fauvel, 1895
1895. Philonthus leptocerus Fauvel, Revue d’Entomologie,
14(6–7): 92–127.
1927.Philonthus pseudooculatus Bernhauer, Arkiv
för Zoologi, 19: 1–28.
Distribution: India (Uttarakhand), Myanmar, Cambodia,
Vietnam, Indonesia (Sumatra).
Philonthus maculatus Cameron, 1920
1920. Philonthus maculatus Cameron, Entomologist’s
Monthly Magazine, 56: 141–148, 214–220.
Distribution: India.
Philonthus malcolmi Herman, 2001
2001. Philonthus malcolmi Herman, Bulletin
of the American Museum of Natural History, 264: 1–82.
1977. Philonthus cameroni Pajni & Kohli, Oriental
Insects, 11(4): 513–519.
Distribution: India (Punjab).
Philonthus modestus Cameron, 1932
1932. Philonthus modestus Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India, Sri Lanka.
Philonthus peliomerus Kraatz, 1859
1859. Philonthus peliomerus Kraatz, Archiv
für Naturgeschichte, 25(1): 1–96.
Distribution: India (Assam, Uttarakhand), Sri Lanka,
Japan, China (Sichuan), Myanmar, Indonesia, French Equatorial Africa, Réunion,
Mauritius, Seychelles, Madagascar, Philippines.
Philonthus perniger Bernhauer, 1920
1920. Philonthus perniger Bernhauer, Archiv
für Naturgeschichte, 84(10): 177–188.
Distribution: India (Tamil Nadu: Madurai).
Philonthus schuhi Schillhammer,
2003
2003. Philonthus schuhi Schillhammer, Koleopterologische
Rundschau, 73: 85–136.
Distribution: India, Nepal, Myanmar, Thailand, Laos,
Vietnam, Malaysia (Penang).
Philonthus suturorubineous Pajni &
Kohli, 1977
1977. Philonthus suturorubineous Pajni &
Kohli, Oriental Insects, 11(4): 513–519.
Distribution: India (Punjab).
Philonthus tamulus Cameron, 1932
1932. Philonthus tamulus Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India.
Genus: Platydracus Thomson, C.G., 1858
1858.Platydracus Thomson,
C. G., Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlingar, 15.
Platydracus (Platydracus)
asemus (Kraatz, 1859)
1859. Platydracus (Platydracus)
asemus (Kraatz), Archiv für Naturgeschichte, 25(1): 1–96.
1911. Staphylinus
aeneicollis Bernhauer, Entomologische Blätter, 7(2–3): 199–200.
Distribution: India
(Meghalaya, Uttar Pradesh, West Bengal, Sikkim: Darjeeling).
Platydracus (Platydracus)
birmanus (Fauvel, 1895)
1895. Platydracus (Platydracus)
birmanus (Fauvel), Revue d’Entomologie, 14(6–7): 180–220.
Distribution: India,
Myanmar, Bhutan.
Platydracus (Platydracus)
brevipennis Smetana & Davies, 2000
2000. Platydracus (Platydracus)
brevipennis Smetana & Davies, American Museum Novitates, 3287:
1–88.
1859.Staphylinus
brachypterus Kraatz, Archiv für Naturgeschichte, 25(1): 1–96.
Distribution: India.
Platydracus (Platydracus)
dudgeoni (Cameron, 1932)
1932. Platydracus (Platydracus)
dudgeoni (Cameron), Taylor & Francis, London. xiii + 443pp.
Distribution: India
(Punjab).
Platydracus (Platydracus)
goryi (Laporte de Castelnau, 1835)
1835. Platydracus (Platydracus)
goryi (Laporte de Castelnau), Méquignon-Marvis Père et Fils, Paris,
159pp.
1859. Staphylinus
auripennis Kraatz, Archiv für Naturgeschichte, 25(1): 1–96.
Distribution: India.
Platydracus (Platydracus)
indicus (Kraatz, 1859)
1859. Platydracus (Platydracus)
indicus (Kraatz), Archiv für Naturgeschichte, 25(1): 1–96.
1859. Ocypus
lineatus Walker, Annals and Magazine of Natural History, 3(3):
370–376.
Distribution: India,
Sri Lanka, Malaysia.
Platydracus (Platydracus)
lomii (Cerruti, 1951)
1951. Platydracus (Platydracus)
lomii (Cerruti), Rivista di Biologia Coloniale, 10: 15–22.
Distribution: India
(Punjab).
Platydracus (Platydracus)
maculipennis (Kraatz, 1859)
1859. Platydracus (Platydracus)
maculipennis (Kraatz), Archiv für Naturgeschichte, 25(1): 1–96.
Distribution: India,
Pakistan, China (Hong Kong).
Platydracus (Platydracus)
marginatus (Cameron, 1944)
1944. Platydracus (Platydracus)
marginatus (Cameron), Proceedings of the Royal Entomological Society of
London (B), 13(1–2): 104–108.
Distribution: India (Sikkim: Darjeeling).
Platydracus (Platydracus)
parvulus Smetana & Davies, 2000
2000. Platydracus (Platydracus)
parvulus Smetana & Davies, American Museum Novitates, 3287:
1–88.
Distribution: India
(Assam).
Platydracus (Platydracus)
semipurpureus (Kraatz, 1859)
1859. Platydracus (Platydracus)
semipurpureus (Kraatz), Archiv für Naturgeschichte, 25(1): 1–96.
1868. Staphylinus
semicupreus Gemminger & Harold, E. H. Gummi, Monachii [Munich].
pp. 425–752.
Distribution: India.
Platydracus (Platydracus)
sparsus (Cameron, 1932)
1932. Platydracus (Platydracus)
sparsus (Cameron), Taylor & Francis, London. xiii + 443pp.
Distribution: India
(Sikkim: Darjeeling).
Platydracus (Platydracus)
submarmorellus (Schubert, K., 1908)
1908. Platydracus (Platydracus)
submarmorellus (Schubert, K.), Deutsche Entomologische Zeitschrift,
(5): 609–625.
Distribution: India
(Himachal Pradesh), Bhutan.
Platydracus (Platydracus)
yolensis (Cerruti, 1951)
1951. Platydracus (Platydracus)
yolensis (Cerruti), Rivista di Biologia Coloniale, 10: 15–22.
Distribution: India
(Punjab).
Platydracus basicornis (Fauvel, 1895)
1895. Platydracus
basicornis (Fauvel), Revue d’Entomologie, 14(6–7): 92–127.
Distribution: India,
Myanmar.
Platydracus bengalensis (Bernhauer, 1914)
1914. Platydracus
bengalensis (Bernhauer), Verhandlungen der Kaiserlich-Königlichen
Zoologisch-Botanischen Gesellschaft in Wien, 64: 76–109.
1914. Staphylinus
bengalensis Bernhauer, Verhandlungen der Kaiserlich-Königlichen
Zoologisch-Botanischen Gesellschaft in Wien, 64: 76–109.
Distribution: India
(West Bengal).
Platydracus curticornis (Fauvel, 1895)
1895. Platydracus
curticornis (Fauvel), Revue d’Entomologie, 14(6–7): 92–127.
Distribution: India,
Myanmar.
Platydracus maculicollis (Fauvel, 1895)
1895. Platydracus
maculicollis (Fauvel), Revue d’Entomologie, 14(6–7): 92–127.
Distribution: India
(Assam, Manipur), Myanmar.
Platydracus purpurascens (Cameron, 1920)
1920. Platydracus
purpurascens (Cameron), Entomologist’s Monthly Magazine, 56: 94–99.
Distribution: India
(Karnataka).
Platydracus virgulatus (Fauvel, 1895)
1895. Platydracus
virgulatus (Fauvel), Revue d’Entomologie, 14(6–7): 92–127.
Distribution: India
(West Bengal), Myanmar.
Genus: Quedius Stephens, 1829
1829. Quedius Stephens,
Baldwin and Cradock, London, 68 pp.
Quedius (Distichalius)
chatterjeei Cameron, 1926
1926. Quedius
(Distichalius) chatterjeei Cameron, Transactions of the Entomological
Society of London, 1925: 341–372.
Distribution: India.
Quedius (Distichalius)
deceptor Cameron, 1944
1944. Quedius
(Distichalius) deceptor Cameron, Proceedings of the Royal Entomological
Society of London (B), 13(1–2): 104–108.
Distribution: India
(Sikkim: Darjeeling, Uttar Pradesh), Nepal.
Quedius (Distichalius)
kashmirensis Cameron, 1944
1944. Quedius (Distichalius)
kashmirensis Cameron, Proceedings of the Royal Entomological Society of
London (B), 13(1–2): 104–108.
Distribution: India
(Kashmir, Uttar Pradesh), Nepal.
Quedius (Distichalius)
taruni Smetana, 1988
1988. Quedius (Distichalius)
taruni Smetana, Quaestiones Entomologicae, 24(2): 163–464.
Distribution: India,
Nepal.
Quedius (Microsaurus)
adjacens Cameron, 1926
1926. Quedius
(Microsaurus) adjacens Cameron, Transactions of the
Entomological Society of London, 1925: 341–372.
Distribution: India
(Kashmir, Himachal Pradesh, Uttar Pradesh), China (Hunan, Sichuan, Shaanxi),
Vietnam.
Quedius (Microsaurus)
antennalis Cameron, 1932
1932. Quedius
(Microsaurus) antennalis Cameron, Taylor & Francis, London.
xiii + 443pp.
Distribution: India
(Sikkim: Darjeeling), Myanmar, China (Fujian, Gansu, Ghizhou, Hainan, Henan,
Hubei, Shaanxi, Sichuan), Laos, Vietnam.
Quedius (Microsaurus)
apicicornis Eppelsheim, 1895
1895. Quedius
(Microsaurus) apicicornis Eppelsheim, Deutsche Entomologische
Zeitschrift, (2): 53–70.
Distribution: India
(Sikkim: Darjeeling), Nepal, Bhutan.
Quedius (Microsaurus)
beesoni Cameron, 1932
1932. Quedius
(Microsaurus) beesoni Cameron, Taylor & Francis, London.
xiii + 443 pp.
Distribution: India
(Sikkim: Darjeeling, Uttar Pradesh), Nepal, China (Chongqing, Fujian, Guangxi,
Guizhou, Hubei, Shaanxi, Shanghai, Sichuan, Yunnan, Zhejiang), Taiwan.
Quedius (Microsaurus)
cruentus (Olivier, A. G., 1795)
1795. Quedius
(Microsaurus) cruentus (Olivier A.G.), Coléoptères, Lanneau,
Paris, 557 pp.
1832. Quedius
erythropterus Stephens, Mandibulata, Baldwin & Cradock, London.
240 pp.
1835. Quedius
analis Stephens, Mandibulata, Baldwin & Cradock, London. pp.
369–477.
1857. Philonthus
putridarius Gistel, Coléoptères, Lanneau, Paris, 557 pp.
1870. Quedius
virens Rottenberg, Berliner Entomologische Zeitschrift, 14(1–2):
11–40.
1913. Quedius
obscurus Lokay, Časopis České Společnosti Entomologické, 10:
136–140.
Distribution: India,
Albania, Austria, Belgium, Bulgaria, Croatia, Czech Republic, Denmark, Finland,
France, Great Britain, Germany, Greece, Hungary, Ireland, Italy, Latvia,
Lithuania, Luxembourg, Netherlands, Norway, Poland, Portugal, Romania, Spain,
Sweden, Switzerland, Ukraine, Armenia, Turkey, Morocco, Algeria, Tunisia,
Massachusetts, Maine, Michigan, New Jersey, New York, Ohio, Pennsylvania.
Quedius (Microsaurus)
dui Smetana, 1988
1988. Quedius
(Microsaurus) dui Smetana, Quaestiones Entomologicae, 24(2):
163–464.
Distribution: India
(Himachal Pradesh).
Quedius (Microsaurus)
flavocaudatus Cameron, 1944
1944. Quedius
(Microsaurus) flavocaudatus Cameron, Proceedings of the Royal
Entomological Society of London (B), 13(1–2): 104–108.
Distribution: India
(Uttar Pradesh), Nepal.
Quedius (Microsaurus)
franzi Smetana, 1975
1975. Quedius
(Microsaurus) franzi Smetana, Oriental Insects, 9(3):
323–342.
Distribution: India
(Sikkim: Darjeeling), Nepal.
Quedius (Microsaurus)
franzi najik Smetana, 1992 (subspecies of Quedius (Microsaurus) franzi.
1992. Quedius
(Microsaurus) franzi najik Smetana, Stuttgarter Beiträge zur Naturkunde
(A: Biologie), 487: 1–11.
Distribution: India
(Sikkim: Darjeeling).
Quedius (Microsaurus)
fulgidus (Fabricius, 1792)
1792. Quedius
(Microsaurus) fulgidus (Fabricius), C. G. Proft, Hafniae,
538pp.
Distribution: India,
Canary Islands, Madeira, Iceland, Libya, Tunisia, Algeria, Turkey, Indonesia,
Peru, Chile, Australia, New Zealand, Chatham Islands.
Quedius (Microsaurus)
inquietus (Champion, 1925)
1925. Quedius
(Microsaurus) inquietus (Champion), Entomologist’s Monthly Magazine,
61: 6–11.
Distribution: India
(Sikkim: Darjeeling, Uttar Pradesh), Nepal, China (Hubei, Shaanxi, Sichuan,
Yunnan), Vietnam.
Quedius (Microsaurus)
milansaar Smetana, 1988
1988. Quedius
(Microsaurus) milansaar Smetana, Quaestiones Entomologicae,
24(2): 163–464.
Distribution: India
(Himachal Pradesh).
Quedius (Microsaurus)
ochripennis (Ménétriés, 1832)
1832. Quedius (Microsaurus)
ochripennis (Ménétriés), Académie Impériale des Sciences, St.
Pétersbourg, 271pp.
1835. Emus floralis Lacordaire, Méquignon-Marvis,
Père et Fils, Paris, 696pp.
1835. Staphylinus laetus Faldermann, Nouveaux
Mémoires de la Société Impériale des Naturalistes de Moscou, 4: 1–310.
1876. Quedius assecla Mulsant & Rey, Annales
de la Société d’Agriculture Histoire Naturelle et Arts Utiles de Lyon,
(4)8: 145–856.
Distribution: India
(Himachal Pradesh), Ukraine, Russia (s European), Armenia, Georgia, Azerbaijan,
Turkey, Syria, Lebanon, Israel, Algeria, Iran, Kazakhstan, Kyrgyzstan,
Tajikistan, Turkmenistan, Uzbekistan, Afghanistan, Pakistan.
Quedius (Microsaurus)
placidus Cameron, 1932
1932. Quedius
(Microsaurus) placidus Cameron, Taylor & Francis, London.
xiii + 443 pp.
Distribution: India
(Sikkim: Darjeeling), Nepal, Bhutan.
Quedius (Microsaurus)
ripicola Cameron, 1926
1926. Quedius
(Microsaurus) ripicola Cameron, Transactions of the
Entomological Society of London, 1925: 341–372.
Distribution: India
(Himachal Pradesh, Sikkim, Uttar Pradesh, Uttarakhand, West Bengal), Nepal.
Quedius (Microsaurus)
spectabilis Kraatz, 1859
1859. Quedius (Microsaurus)
spectabilis Kraatz, Archiv für Naturgeschichte, 25(1):1–96.
Distribution: India.
Quedius (Microsaurus)
stevensi Cameron, 1932
1932. Quedius (Microsaurus) stevensi
Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India (Sikkim, Uttarakhand), Nepal,
China (Yunnan).
Quedius (Raphirus)
anomalus Cameron, 1926
1926. Quedius (Raphirus) anomalus
Cameron, Transactions of the Entomological Society of London, 1925:
341–372.
Distribution: India.
Quedius (Raphirus)
assamensis Cameron, 1932
1932. Quedius (Raphirus) assamensis
Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India (Meghalaya, Nagaland), Nepal,
China (Yunnan).
Quedius (Raphirus)
aureiventris Bernhauer, 1915
1915. Quedius (Raphirus) aureiventris
Bernhauer, Coleopterologische Rundschau, 4(3): 49–60.
Distribution: India (Kashmir, Himachal Pradesh, Uttar
Pradesh, Uttarakhand), Nepal, Bhutan, Pakistan.
Quedius (Raphirus)
daksumensis Coiffait, 1982
1982. Quedius (Raphirus) daksumensis
Coiffait, Entomologica Basiliensia, 7: 231–302.
Distribution: India (Kashmir), Nepal.
Quedius (Raphirus)
durgaa Smetana, 1988
1988. Quedius (Raphirus) durgaa
Smetana, Quaestiones Entomologicae, 24(2): 163–464.
Distribution: India (Sikkim: Darjeeling).
Quedius (Raphirus)
fluviatilis Cameron, 1926
1926. Quedius (Raphirus) fluviatilis
Cameron, Transactions of the Entomological Society of London, 1925: 341–372.
Distribution: India (Kashmir, Himachal Pradesh, Uttar
Pradesh).
Quedius (Raphirus)
himalayicus Bernhauer, 1915
1915. Quedius (Raphirus) himalayicus
Bernhauer, Coleopterologische Rundschau, 4(3): 49–60.
Distribution: India (Sikkim: Darjeeling, Himachal
Pradesh, Nagaland, Uttar Pradesh, Uttarakhand), Nepal.
Quedius (Raphirus)
kaalo Smetana, 1988
1988. Quedius (Raphirus) kaalo
Smetana, Quaestiones Entomologicae, 24(2): 163–464.
Distribution: India (Uttar Pradesh, western Almora),
Nepal.
Quedius (Raphirus)
muscicola Cameron, 1932
1932. Quedius (Raphirus) muscicola
Cameron, Taylor & Francis, London. xiii + 443 pp.
Distribution: India (Sikkim: Darjeeling, Himachal
Pradesh, Uttar Pradesh), Nepal, Myanmar, China (Gansu, Guizhou, Hubei, Shaanxi,
Sichuan, Yunnan), Vietnam.
Quedius (Raphirus)
pahelo Smetana, 1992
1992. Quedius (Raphirus) pahelo
Smetana, Stuttgarter Beiträge zur Naturkunde (A: Biologie), 487: 1–11.
Distribution: India (Sikkim: Darjeeling).
Quedius (Raphirus)
paschim Smetana, 1988
1988. Quedius (Raphirus) paschim
Smetana, Quaestiones Entomologicae, 24(2): 163–464.
Distribution: India (Uttar Pradesh), Nepal.
Quedius (Raphirus)
rugosus Cameron, 1921
1921. Quedius (Raphirus) rugosus
Cameron, Entomologist’s Monthly Magazine, 57: 270–274.
Distribution: India (Sikkim: Darjeeling, Nagaland,
Uttar Pradesh), Nepal, Myanmar.
Quedius (Raphirus)
tonglu Smetana, 1988
1988. Quedius (Raphirus) tonglu
Smetana, Quaestiones Entomologicae, 24(2): 163–464.
Distribution: India (Sikkim: Darjeeling), Nepal.
Quedius aureipilis Cameron, 1932
1932. Quedius (Raphirus) aureipilis Cameron, Taylor
& Francis, London. xiii + 443 pp.
Distribution: India.
Quedius chlorophanus Erichson, 1840
1840. Quedius chlorophanus Erichson, F. H.
Morin, Berlin, pp. 401–954.
Distribution: India (Bengal).
Genus: Staphylinus Linnaeus, 1758
1758. Staphylinus
Linnaeus, Systema naturae Edidito decima, reformata, 824+iiipp.
1886. Ouchemus
Gozis, Imprimerie Herbrin, Montlucon 36pp.
Staphylinus
dimidiaticornis Gemminger, 1851
1851. Staphylinus
dimidiaticornis Gemminger, Ein Beitrag zu den Localfaunen Deutschlands.
Jena: Friedrich Mauke, 65 pp.
1903. Staphylinus
parumtomentosus Stein, G. R., Entomologische Zeitung, 22(4–5): 128.
1913. Staphylinus
ernesti Bernhauer, Entomologische Blätter, 9(9–10): 219–224
1940. Staphylinus
brunneimaculatus Tottenham, Entomologist’s Monthly Magazine, 76:
129–130.
Distribution: India
(Kashmir), Austria, Belgium, Bulgaria, Belarus, Croatia, Czech Republic,
Denmark, Estonia, Finland, France, Great Britain, Germany, Greece, Hungary,
Ireland, Italy, Latvia, Lithuania, Luxembourg, Netherlands, Norway, Poland,
Portugal, Romania, Slovenia, Spain, Sweden, Switzerland, Ukraine.
Tribe: Xantholinini Erichson, 1839
1837. Gyrohypnidae
Kirby, Fauna Boreali-Americana, xxxix + 325 + [1]pp.
1839. Xantholinini
Erichson, Genera et species staphylinorum insectorum coleopterorum familiae,
F. H. Morin, Berlin, 400 pp.
1957. Gyrohypnini
Hatch, University of Washington Publications in Biology, 16 (2).
Genus: Indolinus Bordoni, 2002
2002. Indolinus Bordoni,
Monografie, Museo Regionale di Scienze Naturali, Torino, 33.
Indolinus mitomorphoides (Coiffait,
1984)
1984. Xantholinus
mitomorphoides Coiffait, Annales de la Société Entomologique de
France, 20(4): 373–387.
Distribution: India
(Meghalaya, Uttarakhand, West Bengal), Nepal, Bhutan, Myanmar, Thailand, Laos.
Genus: Indomorphus Bordoni, 2002
2002. Indomorphus Bordoni,
Monografie, Museo Regionale di Scienze Naturali, Torino, 33.
Indomorphus parcus (Eppelsheim,
1895)
1895. Xantholinus
parcus Eppelsheim, Deutsche Entomologische Zeitschrift, (2): 385–408.
1982. Xantholinus
bhutanicus Coiffait, Entomologica Basiliensia, 7: 231–302.
Distribution: India
(Uttar Pradesh), Nepal, Bhutan, Myanmar, China (Sichuan, Yunnan).
Indomorphus shavrini Bordoni, 2013
2013. Indomorphus
shavrini Bordoni, Fragmenta Entomologica, 45(1–2): 59–63.
Distribution: India
(Uttarakhand).
Thus, a total of 199
species belonging to 13 genera of subfamily Staphylininae have been reported
out of which Staphylinini is the most speciose with 93% of the known species,
followed by Othiini and Xantholinini (Figure 1). Moreover, 73 species belonging
to genera Othius (6), Bisnius (2), Heterothops (4),
Neobisnus (1), Philonthus (36), Platydracus (12), Quedius
(11), Indomorphus (1) have only been reported from India constituting
around 36.68% of the catalogued rove beetle diversity. At generic level, Philonthus,
a lineage of specialized predators, is the most diverse group of rove beetles
with 105 species followed by Quedius (36) and Platydracus (20)
(Figure 2). Holarctic genus Quedius is mainly restricted to higher
altitude regions of India namely in the states of Kashmir, Himachal Pradesh,
Manipur, Meghalaya, Sikkim, Uttarakhand and West Bengal. Platydracus, on
the other hand, is a group of large rove beetles, which prefer to predate on
other insects feeding on the dead decaying matter. All the three genera
together constitute 80% of the recorded diversity so far.
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References
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Al-Mekhlafi (2023). Beetles associated with buried carcasses:
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Sciences 30(7): 1–8. https://doi.org/10.1016%2Fj.sjbs.2023.103706
Assing, V. (2008). A revision of
Othiini. XVI. Four new species of Othius from the Himalaya and China,
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