Journal of Threatened
Taxa | www.threatenedtaxa.org | 26 June 2024 | 16(6): 25461–25464
ISSN 0974-7907
(Online) | ISSN 0974-7893 (Print)
https://doi.org/10.11609/jott.8988.16.6.25461-25464
#8988 | Received 25 February 2024 | Final received 19 May 2024 | Finally
accepted 12 June 2024
First record of Croaking Gourami Trichopsis vittata (Cuvier,
1831) from West Bengal, India
Sujal Dutta 1 , Bakul
Biswas 2 & Bibhas Guha 3
1–3 Department of Zoology, Netaji
Subhas Open University, DD-26, Sector-I, Salt Lake City, Kolkata, West Bengal
700064, India.
1 sujalsnehadutta@gmail.com, 2
bakul87in@gmail.com, 3 g.bibhas@gmail.com (corresponding
author)
Editor: J.A. Johnson, Wildlife Institute of India,
Dehradun, India. Date of publication: 26 June 2024
(online & print)
Citation: Dutta,
S., B. Biswas & B. Guha (2024). First record of Croaking Gourami Trichopsis vittata
(Cuvier, 1831) from West Bengal, India. Journal of Threatened Taxa 16(6): 25461–25464. https://doi.org/10.11609/jott.8988.16.6.25461-25464
Copyright: © Dutta et al. 2024. Creative Commons Attribution 4.0 International
License. JoTT
allows unrestricted use, reproduction, and distribution of this article in any
medium by providing adequate credit to the author(s) and the source of
publication.
Funding: Self-funded.
Competing interests: The authors declare no competing interests.
Acknowledgements: The authors are thankful to the
Department of Zoology, Netaji Subhas Open University for providing requisite
research facilities. The authors are also thankful to Fresh water fish section, Zoological Survey of India, Kolkata for proper species
identification and documentation.
Abstract: The Croaking Gourami Trichopsis vittata (Cuvier,
1831) is a native fish to southeastern Asia and Sundaland,
with introduction reports from the USA, Philippines, and many other countries
including India. In India, this species was first reported from Chembarampakkam Lake situated in Chennai during the year
2015. This study reports the presence of this fish for the first time in the Magra Beel, a wetland in the
district of Nadia, West Bengal, in 2021, 2022, 2023. The laboratory evaluated
the fish samples taken from the marsh to determine their fin ray counts and
morphometric data. The results identified the species as T. vittata and supported previous research.
Keywords: Aquarium trade, distribution,
invasion, Magra Beel,
meristic count, morphometry, osphronemid fish,
species identification.
Trichopsis vittata (Cuvier,
1831), known as Croaking Gourami in the ornamental fish trade, is a small
air-breathing osphronemid fish, reported as native
from Java, Borneo, Sumatra, Peninsular Malaysia, Thailand, and the Mekong basin
in Cambodia, Laos, and Viet-nam (Kottelat
1985; Baird et al. 1999). The species is an obligate air-breathing fish that
prefers to live in shallow sluggish or standing-water habitats with abundant
vegetation (Rainboth 1996). It can survive in
brackish water up to 20 ppt salinity, and in temperatures down to 7.20C
(Schofield & Schulte 2016). It feeds on small planktonic crustaceans and
insect larvae (Rainboth 1996). This species comprises
more-extended filamentous anal-fin rays extending almost to the tip of the
caudal-fin and also possesses three dark lateral stripes on the body. A typical
dark spot is present above the pectoral fin of the fish as a unique identifying
character. Unlike other osphronemid fishes, T. vittata is popularized for the production of audible
sounds or “croaks” (Ladich & Yan 1998).
Trichopsis vittata has shown invasiveness and introduced itself to
a larger extent. Since its description by Cuvier in 1831 it has extended its
range. The feral population has been documented in the USA (Florida) and the
Philippines (Shafland 1996; Schofield & Pecora
2013; Schofield & Schulte 2016). The southeastern Asian countries including
Indonesia, Thailand, Java, Borneo, Sumatra (Rainboth
1996; Kottelat 1985; Liengpornpan
et al. 2007; Beamish et al. 2010), Vietnum (Freyhof et al. 2000; Herder & Freyhof
2006), Mekong tributaries of Indochina (Robins 1992; Baird et al. 1999; Kottelat 2013), Laos (Martin et al. 2011), Singapore
(Alfred 1996; Low & Lim 2012), Cambodia (Rainboth 1996), Malaysia (Beamish et al. 2003) are reported
to be the native places of T. vittata.
The non-native population of T. vittata has
been documented from Myanmar and Bangladesh (Noren et
al. 2017; Akash et al. 2018; Hossain et al. 2019). Literature depicts that in
India T. vittata was first reported from the Chembarampakkam Lake situated in southern (Knight &
Balasubramanian 2015); but its existence has not yet been reported from West
Bengal.
Study
Area and Methods
The fish samples were first collected on 1 August 2021 from the Magra Beel. The collection was
done by using a hand held drag net as per the conventional method. Then in
quest of other populations of fish, multiple field visits were done between
August 2021 and July 2023 in different fishing points of the Magra Beel within a distance of
1.5 km (Point-1: Opposite to Bijra Primary School;
Point-2: Concrete bridge at 3 number Magra Beel; Point-3: Kheya Ghat opposite to Birahi Union Tapashili Fishermen’s Cooperative Society Limited office).
The sampling sites of the Magra Beel
are presented in Image 1.
After collection, the fish specimens were immediately preserved in 10%
formalin for taxonomic study (Joshi et al. 2015). Different morphometric
characteristics including total length, standard length, pectoral fin base
length, pelvic fin base length, and anal fin base length were measured as per
the conventional method and different meristic counts for dorsal fin rays, anal
fin rays, pectoral fin rays, pelvic fin rays were observed accordingly (Murdy & Shibkawa 2001). The
preserved fish samples were submitted to the Fresh Water Fish Section of the
Zoological Survey of India, Kolkata for identification and documentation. Some
fish samples were subsequently transferred into 30%, 50%, and 70% ethanol for
long-term preservation (Sterba 1962; Talwar & Jhingran 1991) for further study.
Results
Morphometric
and meristic characters
The
collected samples have laterally elongated bodies with a standard length of
4.10 cm (Table 1). The dorsal fin originates far behind the base of the pectoral
fin corroborating the previous findings of Hossain et al. (2019) and Shefat et al. (2020). A thin black line is found below the
eye (Noren et al. 2017). Stripes and black spots are
visible on the fin of both sides of the body. The morphometric measurements
including total length, standard length, pectoral fin base length, pelvic fin
base length, and anal fin base length of the fish have been presented in Table
1. The anal fin is comprised of multi-branched fin rays with 6–8 spines and a
few elongated filaments-like rays which are extended almost to the tip of the
caudal fin. The dorsal fin of T. vittata has
2–4 spines, while the pelvic fin contains one spine followed by a filament and
four rays. On the posterior half of the body, there are three longitudinal dark
bands, which help to distinguish T. vittata
from the related species T. schalleri and T.
pumila (Noren et al.
2017). The fin ray counts are detailed in Table 1 and are consistent with the
findings reported by Hossain et al. (2019).
Identification
and cataloguing
After
collecting the samples of fish from three different spots, the specimens were
submitted to the Zoological Survey of India (ZSI) for identification and
cataloguing. The ZSI has identified the species and registered the submitted
specimens as Trichopsis vittata
(Cuvier, 1831) [Regd. No. ZSI FF 9935 dt. 24. 08.
2023].
Discussion
In this
present study, T. vittata was found in a lower
abundance than the other native fish species; but the overall count is
sufficient to prove its strong adaptability to this new environment. T. vittata is an efficient invader that can survive in a
small volume of water and can tolerate high pollution and low dissolved oxygen
level conditions (Wongsiri, 1982). Knight & Balasubramanium (2015) reported that in India T. vittata may compete for niche space with native species
such as Trichogaster lalius
(Hamilton, 1822) and Spiketail Paradise Fish Pseudosphromenus cupnus
(Cuvier, 1831). The negative effects of T. vittata
may also include aggressive displacement of native species or it may also act
as a vector for parasites or pathogens such as trematode, Euclinostomum
heterostomum (Purivirojkul
& Sumontha 2013). Thus, the spreading of T. vittata across the country over the years will affect
the native fish biodiversity. Of late, the particular impact of this species on
other indigenous fish species is currently unknown, therefore, further in-depth
study regarding its biology, ecology, and inter-specific interactions among the
non-native range in India are required at the earnest level.
The way of
introduction of the fish species in the Magra Beel is not known. However, the most probable mechanism of
introduction may be the release from the aquarium trade (Noren
et al. 2017). As per the first recommendations of ZSI, Kolkata, “is an exotic
species and distributed worldwide via aquarium trade. The specimen shows some
variations from the description of Trichopsis
vittata and may be a hybrid also. It needs
further study based on freshly collected specimens for confirmation”. Later on ZSI Kolkata identified and registered the specimen as Trichopsis vittata
(Cuvier, 1831), family Osphronemidae [Regd No. ZSI FF 9935 dt. 24. 08.
2023]. Deliberate aquaculture efforts could serve as a vector for invasion
(Akash & Hossain 2018). Given that T. vittata
has been previously reported in various rivers across Bangladesh (Noren et al. 2017), some of which are in close proximity to
the state of West Bengal, it is plausible that this species could migrate to
West Bengal via these river routes.
Table 1. The
morphometric measurement and meristic count of fin rays in Trichopsis
vittata.
|
Morphometric characters |
Length (cm) |
Meristic characters |
Number (range) |
|
Total length |
5.70 |
Dorsal fin ray |
7–8 |
|
Standard length |
4.10 |
Pectoral fin ray |
10–11 |
|
Pectoral fin base length |
1.20 |
Ventral fin ray |
7–9 |
|
Pelvic fin base length |
2.30 |
Anal fin ray |
31–33 |
|
Anal fish base length |
1.60 |
Caudal fin ray |
12–16 |
|
- |
- |
Lateral line scales |
29 |
For images
- - click here for full PDF
References
Akash, M. & M.A. Hossain (2018). A Southeast
Asian species in the Ganges delta: on spreading extent of non-native croaking
gourami Trichopsis vittata
(Cuvier, 1831) in Bangladesh. Bio-invasion Record 7: 447–450. https://doi.org/10.3391/bir.2018.7.4.16
Alfred, E.R. (1966). The fresh-water fishes of Singapore. Zoologische Verhandelingen
78: 3– 68.
Baird, I.G., V. Inthaphaisy, P. Kisouvannalath, B. Phylavanh
& B. Mounsouphom (1999). The Fishes
of Southern Lao. Lao Community Fisheries and Dolphin Protection Project. Ministry of
Agriculture and Forestry, Lao PDR, 161 pp.
Beamish, F.W.H., R.B. Beamish & S.L.H. Lim (2003). Fish
Assemblages and Habitat in a Malaysian Blackwater Peat Swamp. Environmental
Biology of Fishes 68: 1–13. https://doi.org/10.1023/A:1026004315978
Beamish, F.W.H., P. Plongsesthee, P. Chanintarapoomi & P. Nithirojpakdee
(2010). Total length-weight relationships among Thai freshwater fishes and the
influence of capture location and preservation. Journal of Applied
Ichthyology 27(3): 955–958. https://doi.org/10.1111/j.1439-0426.2010.01620.x
Cuvier, G. & A. Valenciennes (1831). Histoire naturelle des poissons. Tome 7.
Paris, F.G. Levrault, 531 pp. https://doi.org/10.5962/bhl.title.12283
Freyhof, J., D.V. Serov
& T.N. Nguyen (2000). A preliminary checklist of the freshwater
fishes of the River Dong Nai, South Vietnam. Bonner
Zoologische Beiträge
49: 93–99.
Hamilton, F. (1822). An account of the fishes found in the river
Ganges and its branches. Edinburgh & London. pp. 1–39. https://doi.org/10.5962/bhl.title.6897
Herder, F. & J. Freyhof (2006). Resource
partitioning in a tropical stream fish assemblage. Journal of Fish Biology
69(2): 571–589. https://doi.org/10.1111/j.1095-8649.2006.01126.x
Hossain, M.D., S. Shoaibe, L. Gulshan & M.D. Obaida (2019). First
record of invasive croaking gourami, Trichopsis
vittata (Cuvier, 1831) in South-eastern
Bangladesh. Journal of Fisheries and Life Sciences 4(2): 6–9.
Joshi, K.K., M.S. Varsha, V.L. Sruthy, V.
Susan & P.V. Prathyusha (2015). Basics of
sample collection, preservation and species identification of finfish. Summer
school on Recent Advances in Marine Biodiversity Conservation and Management,
CMFRI.
Knight, J.M. & S. Balasubramanian (2015). On a record
of two alien fish species (Teleostei: Osphronemidae) from the natural waters of Chennai, Tamil
Nadu, India. Journal of Threatened Taxa 7(3): 7044–7046. https://doi.org/10.11609/JoTT.o4135.7044-6
Kottelat, M. (1985).
Fresh-water fishes of Kampuchea. Hydrobiologia
121: 249–279.
Kottelat, M. (2013). The fishes
of the inland waters of Southeast Asia: a catalogue and core bibliography of
the fishes known to occur in freshwaters, mangroves and estuaries. Raffles
Bulletin of Zoology 27: 1–663
Ladich, F. &
H.Y. Yan (1998). Correlation between auditory sensitivity and vocalization in anabantoid fishes. Journal of Comparative Physiology A
182: 737–746. https://doi.org/10.1007/s003590050218
Liengpornpan, S., M. Jaroensutasinee & K. Jaroensutasinee
(2007). Biology of croaking gourami, Trichopsisvittata:
the fish that croaks. Thaksin University Journal 10: 72–83.
Low, B.W. & K.K.P. Lim (2012). Gouramies of the genus Trichopodus
in Singapore (Actinopterygii: Perciformes: Osphrenemidae). Nature in Singapore 5: 83–93.
Martin, S.M., K. Lorenzen, R.I. Arthur, P. Kaisone
& K. Souvannalangsy (2011). Impacts of
fishing by dewatering on fish assemblages of tropical floodplain wetlands: a
matter of frequency and context. Biological Conservation 144(1):
633–640. https://doi.org/10.1016/j.biocon.2010.11.005
Murdy, E.O. &
K. Shibukawa (2001). A revision
of the gobiid fish genus Odontamblyopus
(Gobiidae: Amblyopinae).
Ichthyological Research 48: 31–43. https://doi.org/10.1007/s10228-001-8114-9
Norén, M., S. Kullander, M.D.
Rahman & A. Mollah (2017). First
records of Croaking Gourami, Trichopsis vittata (Cuvier, 1831) (Teleostei:
Osphronemidae), from Myanmar and Bangladesh. Check
List 13: 81–85. https://doi.org/10.15560/13.4.81
Purivirojkul, W. &
M. Sumontha (2013). Eucliinostomum heterostomum (Rudolphi,
1809) metacercarial infection in three osphronemid
fish species. Walailak Journal of Sciences
and Technology 10(1): 97–102. https://doi.org/10.2004/wjst.v10i1.423
Rainboth, W.J.
(1996). FAO species identification field guide for fishery purposes. Fishes of
the Cambodian Mekong. Food & Agriculture Org., Fisheries, 265 pp. https://www.fao.org/tempref/docrep/fao/010/v8731e/v8731e.zip
Robins, C.R. (1992). Common and scientific names of fishes from
the United States and Canada. American Fisheries Society (USA) Special
Publication#20, 243 pp. https://doi.org/10.47886/9781934874318
Schofield, P.J. & D.J. Pecora (2013). Croaking
gourami, Trichopsis vittata
(Cuvier, 1831), in Florida, USA. Bio-Invasions Records 2: 247–251.
Schofield, P.J. & J.M. Schulte (2016). Small but
tough: What can ecophysiology of croaking gourami Trichopsis
vittata (Cuvier, 1831) tell us about invasiveness
of non-native fishes in Florida? NeoBiota 28:
51. https://doi.org/10.3897/neobiota.28.5259
Shafland, P.L.
(1996). Exotic fishes of Florida - 1994. Reviews in Fisheries Science
4: 101–122.
Shoaibe, S. &
M.D. Hossain (2020). Distribution and Taxonomic Study of a Newly
Recorded Croaking Gourami, Trichopsis vittata (Cuvier, 1831) in Bangladesh. Acta
Scientific Agriculture 4(1): 107–111. http://doi.org/10.31080/ASAG.2020.04.754
Sterba, G. (1962). Freshwater
Fishes of The World. Vista Books, London, 878 pp. http://doi.org/10.1017/S0030605300002532
Talwar, P.K. & A.G. Jhingran (1991). Inland
Fishes of India and Adjacent Countries. Oxford-IBH Publishing Co. Pvt. Ltd.,
New Delhi, 1158 pp. https://doi.org/10.1007/BF00043269
Wongsiri, S. (1982).
Preliminary survey of the natural enemies of mosquitoes in Thailand. Journal
of the Science Society of Thailand 8: 205–213. https://doi.org/10.2306/scienceasia1513-1874.1982.08.205