Journal of Threatened
Taxa | www.threatenedtaxa.org | 26 November 2023 | 15(11): 24284–24287
ISSN 0974-7907
(Online) | ISSN 0974-7893 (Print)
https://doi.org/10.11609/jott.8683.15.11.24284-24287
#8683 | Received 10 August 2023 | Final received 01 October 2023 |
Finally accepted 31 October 2023
First photographic record of
Hoary-bellied Squirrel Callosciurus pygerythrus (I. Geoffroy Saint Hilaire, 1832) (Mammalia:
Rodentia: Sciuridae) from Banke
National Park, Nepal
Yam Bahadur Rawat 1, Shyam Kumar Shah 2, Sunjeep
Pun 3 & Dristee Chad 4
1,2 Department of National Parks and
Wildlife Conservation, PO Box : 860, Babarmahal, Kathmandu, Nepal.
3 Zoological Society of London
Nepal Office, Po Box 5867, Kathmandu, Nepal.
4 Tribhuvan University, Institute
of Forestry, PO
Box: 43, Hariyokharka-15, Pokhara,
Nepal.
1 yam.rawat@nepal.gov.np
(corresponding author), 2 shyamkumar_shah@yahoo.com, 3 sunjeep.pun@zsl.org,
4 dristim18@gmail.com
Editor: Murali Krishna Chatakonda,
Amity University, Noida, India. Date of publication: 26 November
2023 (online & print)
Citation: Rawat,
Y.B., S.K. Shah, S. Pun & D. Chad (2023). First photographic record
of Hoary-bellied Squirrel Callosciurus pygerythrus (I. Geoffroy Saint Hilaire, 1832)
(Mammalia: Rodentia: Sciuridae) from Banke National Park, Nepal. Journal of Threatened Taxa 15(11): 24284–24287. https://doi.org/10.11609/jott.8683.15.11.24284-24287
Copyright: © Rawat et al. 2023. Creative Commons Attribution 4.0 International
License. JoTT
allows unrestricted use, reproduction, and distribution of this article in any
medium by providing adequate credit to the author(s) and the source of
publication.
Funding: None.
Competing interests: The authors declare no competing interests.
Acknowledgements: We are very grateful to the
Department of National Parks and Wildlife Conservation, Nepal, the Nepal Army Arjunban Battalion, ZSL Nepal, the joint patrol team of
both Banke National Park and the Nepal Army team
deployed in the anti-poaching operation, Senior Game Scout Deepak Chaulagain, Game Scout Binod Malla,
and Game Scout Chakra Shahi. We are also thankful to assistant conservation
officer Mr. Uttam Kumar Chaudhary for his continuous
encouragement and guidance to the team during the operation.
The southern and southeastern
Asia’s forest ecosystems are home to a high diversity and endemism of squirrels
(Koprowski & Nandini 2008; Krishna et al. 2016).
There are 285 species of squirrels worldwide (Thorington
et al. 2012), however, only 12 species including the Hoary-bellied Squirrel
have been recorded from Nepal (Thapa 2014). Squirrels contribute significantly
to ecosystem services by eating fungal spores, seeds, and other plant material,
and they are also important prey for a variety of species in temperate and
tropical forests (Koprowski & Nandini 2008). They
are primarily forest-dependent creatures that can only adapt to a certain level
of habitat loss. Anthropogenic impacts on both small and large mammals in Nepal
appear to be increasing threats to the country’s wildlife, which is leading to
a decline in species numbers (Baral & Shah 2008;
Bhandari & Chalise 2016; Bhandari et al. 2020). A
large number of rodents and lagomorphs also experience this as a result of the
prolonged usage of pesticides on agricultural grounds (Baral
& Shah 2008; Aktar et al. 2009). The
Hoary-bellied Squirrel dwells in patches of dense to moderately dense evergreen
forests in mid-canopy temperate, tropical, and subtropical moist habitats (Molur et al. 2005). The squirrel has been found in
Nepal’s riverine woodland, mixed broad-leaf forest zone, central and eastern
Siwalik foothills, Mahabharat range (Mitchell 1979), as well as modified and
altered habitats including settlements and farmlands. This species, being
arboreal and diurnal, consumes seeds, nuts, fruits, buds, flowers, and insects
(Mitchell 1979) and gathers water from young twigs and bamboo (Karki 2013).
The Hoary-bellied Squirrel is a
widely distributed species in southern Asia (Thorington
et al. 2012; Karki 2013). It can be found in Nepal’s lowlands (300 m or less)
and temperate regions (up to 2,500 m) (Baral &
Shah 2008; Thapa et al. 2016), Sal forests to mixed broad leaved forests in
Mahabharat range (Thapa et al. 2016). The species is distributed throughout the
Siwalik (Chure) and Mahabharat ranges (Jnawali et al. 2011); confirmed from several districts and
protected areas of Nepal (Thapa et al. 2016) (Figure 1). The main threats to
the species in Nepal are habitat loss and hunting for food and medicine (Jnawali et al. 2011). Based on an array of studies,
anthropogenic pressure, habitat degradation, hunting, and pesticide usage are
the major threats to the species (Shrestha 1997; Majpuria
& Majupuria 2006; Baral
& Shah 2008; Thapa et al. 2016). It is classified as a ‘Least Concern’
species globally as well as nationally by the IUCN Red List of Threatened
Species (Jnawali et al. 2011; Duckworth 2016). The
species has yet to be documented from the western part of the country.
Banke National Park (BaNP) established as the 10th national park of
Nepal on 12 July 2010 is situated in the western part of the country. The core
area of the park is entirely located in Banke
District, Lumbini Province, while some portion of the buffer zone area also
lies in Salyan District, Karnali
Province. BaNP is linked with a trans-boundary
landscape, joining Suhelwa Wildlife Sanctuary in
India through Kamdi corridor including national and
community forests towards the south and Bardiya
National Park (BNP) towards the west, which further links with Katerniaghat Wildlife Sanctuary in India via the Khata corridor, national forest, and community forests. The
park consists of eight vegetation types: Sal Shorea
robusta forest, deciduous riverine forest,
savannahs & grasslands, mixed hardwood forest, flood plain, Bhabar, and Chure Range
foothills. These ecosystems harbour 34 species of
mammals, 236 species of birds, nine species of amphibians, 24 species of
reptiles, and 55 species of fish (BaNP 2018). About
90 percent of natural forest coverage is composed of mainly Sal Shorea robusta,
Karma Terminalia tomentosa, Khair Senegalia catechu,
and Sissoo Dalbergia
sissoo species that support a wide variety of
wildlife. Among rodents, only four species, viz., Northern Palm Squirrel Funambulus pennantii,
Indian Crested Porcupine Hystrix indica, Red Giant Gliding Squirrel Petaurista
petaurista, and House Rat Rattus rattus have been previously recorded in the park (BaNP 2018).
The Chure
Hills cover the majority of Banke National Park,
along with some flat areas. It is difficult to monitor wildlife crime and
forest product smuggling in such locations with regular patrols. On 18 June
2023, a team of 10 persons from the BaNP and the
Nepal Army were dispatched for a four-day camping operation. On the first day,
the team walked from the Mahadeva area to Baghsal Sota (also known as Baghsal Khola), a perennial source of water. At 1540 h, while
heading towards Chure Top along Gangate
Khola (a tributary of Baghsal
Sota), the first author noticed an animal climbing on
a Jamun Syzygium cumini
tree at 28.235701° N, 81.826066° E, at an elevation of 532 m (Figure 2). When
the species was approached at close proximity, it leaped to the Bhorla Bauhinia vahlii, a
climber and reached the canopy of the Jamun tree. The species was
photographed with a Canon 2000D, and the location was marked with a Garmin 64s
GPS. Binoculars (Olympus 8 x 42) were used to observe the species size,
coloration, physical traits, and climbing and resting events in the field.
The animal has brown dorsal
pelage but dark grey at limbs and in addition a yellowish hip patch, which
occurs seasonally a blunt muzzle and comparatively larger tail, dirty white
pelage, and alternating bands of black to blackish and yellowish to white
appears in the tail. Based on the characteristics, the animal was identified as
the juvenile Hoary-bellied Squirrel Callosciurus pygerythrus (Image 1 & 2).
The current photographic record
provides the first photographic evidence and range extension of the
Hoary-bellied Squirrel’s distribution in the western part of the country. In
addition, there is a lack of information regarding their population ecology,
interactions with other sympatric rodents like the Orange-bellied Himalayan
Squirrel Dremomys lokriah,
and how the species react to an increase in anthropogenic interactions in
human-dominated landscapes. However, an in-depth study of habitat
characteristics, nutrition, distribution, movement, and activity patterns would
further aid in the long-term survival of Hoary-bellied Squirrel in BaNP.
For
figures & images - - click here for full PDF
References
Aktar, M.W., D. Sengupta & A.
Chaudhary (2009). Impact of pesticides use in agriculture: their benefits and hazards. Interdisciplinary
Toxicology 2(1): 1–12. https://doi.org/10.2478/v10102-009-0001-7
BaNP (2018). Banke
National Park and its Buffer Zone Management Plan 2075/76-2079/80, Banke National Park Office, Ohbari,
Banke Nepal.
https://dnpwc.gov.np/media/publication/Banke_National_Park_Management_Plan_pdf.
Baral, H.S. & K.B. Shah (2008). Wild Mammals of Nepal.
Himalayan Nature, Kathmandu, 158 pp.
Bhandari, S.
& M.K. Chalise (2016). People’s attitudes toward
Striped Hyaena (Hyaena hyaena) Linnaeus, 1758) (Mammalia: Carnivora: Hyaenidae) conservation in lowland Nepal. Journal
of Threatened Taxa 8(9): 9125–9130. https://doi.org/10.11609/jott.2518.8.9.9125-9130
Bhandari, S.,
C. Morley, A. Aryal & U.B. Shrestha (2020). The diet of the striped hyena in
Nepal’s lowland regions. Ecology and Evolution 10: 7953–7962. https://doi.org/10.1002/ece3.6223
Duckworth,
J.W. (2016). Callosciurus pygerythrus. The IUCN Red List of Threatened
Species 2016:e.T3604A22253451. https://doi.org/10.2305/IUCN.UK.20162.RLTS.T3604A22253451.en. Accessed
on 05 September 2023.
Jnawali, S.R., H.S. Baral,
S. Lee, K.P. Acharya, G.P. Upadhyay, M. Pandey, R. Shrestha, D. Joshi, B.R. Lamichhane, J. Griffiths, A.P. Khatiwada,
N. Subedi & R. Admin (Compilers) (2011). The Status of Nepal Mammals: The
National Red List Series, Department of National Parks and Wildlife
Conservation, Kathmandu, Nepal, vii + 266 pp.
Karki, R.
(2013). Distribution
and behavioral ecology of Irrawaddy squirrel (Callosciurus
pygerythrus) in urban habitats of Bhaktapur.
M.Sc. Thesis. Central Department of Environmental Science, Tribhuvan University
Kirtipur, Kathmandu, Nepal, 56 pp.
Koprowski, J.L. & R. Nandini
(2008). Global hotspots and knowledge
gaps for tree and flying squirrels. Current Science 95(7): 851–856.
Krishna,
M.C., A. Kumar, O.P. Tripathi & J.L. Koprowski
(2016). Diversity,
distribution and status of gliding squirrels in protected and non-protected
areas of eastern Himalayas in India. Hystrix 27(2):
111–119. https://doi.org/10.4404/hystrix-27.2-11688
Majupuria, T.C. & R.K. Majupuria (2006). Wildlife and protected areas
of Nepal [Resource and Management]. S. Devi. Saharanpur, India, 427 pp.
Mitchell,
R.M. (1979). The sciurid
rodents (Rodentia: Sciuridae) of Nepal. Journal of
Asian Ecology 1: 21–28.
Molur, S., C. Srinivasulu,
B. Srinivasulu, S. Walker, P.O. Nameer
& L. Ravikumar (2005). Status of South Asian Non-volant Small Mammals: Conservation Assessment
and Management Plan (C.A.M.P) Workshop Report. Zoo Outreach
Organization/CBSG-South Asia, Coimbatore, India, 618 pp.
Shrestha,
T.K. (1997). Mammals
of Nepal with Reference to those of India, Bangladesh, Bhutan and Pakistan.
Mrs. Bimala Shrestha, Kathmandu, Nepal, 371 pp.
Thapa, S.
(2014). A checklist
of mammals of Nepal. Journal of Threatened Taxa 6(8):
6061–6072. https://doi.org/10.11609/JoTT.o3511.6061-72
Thapa, S.,
H.B. Katuwal, S. Koirala, B.V. Dahal,
B. Devkota, R. Rana, H. Dhakal
& H. Basnet (2016). Sciuridae (order: Rodentia) in
Nepal. Small Mammals Conservation and Research Foundation: Kathmandu,
Nepal, 70 pp.
Thorington, R.W., J.L. Koprowski,
M.A. Steele & J.F. Whatton (2012). Squirrels of the world.
Johns Hopkins University Press, Baltimore, Maryland, 459 pp.