Journal of Threatened
Taxa | www.threatenedtaxa.org | 26 February 2025 | 17(2): 26515–26529
ISSN 0974-7907 (Online)
| ISSN 0974-7893 (Print)
https://doi.org/10.11609/jott.8254.17.2.26515-26529
#8254 | Received 03
November 2022 | Final received 28 January 2025 | Finally accepted 05 February
2025
A study on the
diversity of butterflies in selected landscapes of the Indian Institute of
Technology, Guwahati campus, Assam, India
Uma Dutta 1,
Sonali Dey 2 & Deepshikha
Moran 3
1,2 Cell and Molecular
Biology Laboratory, Department of Zoology, Cotton University, Guwahati, Assam
781001, India.
3 Department of
Zoology, Dibru College, Dibrugarh, Assam 786003, India.
1 umadutta@yahoo.com (corresponding author), 2 sonalidey167@gmail.com,
3 deepshikhamoran6@gmail.com
Editor: Jatishwor Singh Irungbam, Centrum Algatech,
Třeboň, Česká Republika. Date of publication: 26 February
2025 (online & print)
Citation: Dutta, U., S. Dey & D. Moran (2025). A study
on the diversity of butterflies in selected landscapes of the Indian Institute
of Technology, Guwahati campus, Assam, India. Journal of Threatened Taxa 17(2): 26515–26529. https://doi.org/10.11609/jott.8254.17.2.26515-26529
Copyright: © Dutta et al. 2025. Creative Commons Attribution 4.0 International License. JoTT allows
unrestricted use, reproduction, and distribution of this article in any medium
by providing adequate credit to the author(s) and the source of publication.
Funding: This research did not receive any specific grant
from funding agencies in the public, commercial, or not-for-profit sectors.
Competing interests: The authors declare no competing interests.
Author details: Dr. Uma Dutta—an associate professor at the
Department of Zoology, Cotton University. She is the
former Dean of Life Sciences and Head of the Department of Zoology, CU. Her
research focuses on bio-active components in chronic disease treatment viz.,
cancer, diabetes cardiovascular disease as well as validation of traditional knowledge and its practices in agricultural pest control
with ethnopharmacological importance. She is passionate about nature and
biodiversity, especially insect, butterfly and birds. She has also published
book on butterfly diversity of Cotton University campus; Sonali Dey–a UGC-JRF PhD research scholar at
Department of Zoology, Cotton University; Dr. Deepshikha Moran–assistant professor, Department of
Zoology, Dibru College, Dibrugarh, she has a good number of publications in
journals of national and international reputes.
Author contributions: UD—contributed to the study design, conceptualisation, supervision,
draft writing, review editing, data collection, spotting of the species and
photography. SD—contributed to data collection, survey
work, photography, draft preparation, figure and table preparation and
bibliographic study. DM—contributed to
table and figure preparation and some part of survey work. The first draft of
the manuscript was written by Uma Dutta and edited by Sonali Dey. All authors commented
on previous versions of the manuscript. All authors read and approved the final
manuscript.
Acknowledgements: Authors are highly grateful to the
workers and residents of IIT Guwahati campus for their
help and support in finding the places in and around the institute. Authors are
also thankful to Miss Bhanita Bora for her help during preparation of the
article.
Abstract: A study has been
carried out to find out the diversity of butterflies at the Indian Institute of
Technology, Guwahati campus, Assam India from September 2019 to March 2022. In
the present study, a total of 82 species with a total of 1,378 individuals of butterflies
belonging to six families, namely, Papilionidae, Pieridae, Lycaenidae,
Nymphalidae, Hesperiidae, and Riodinidae have been recorded. During the survey,
the maximum number of butterflies were observed in the old E-type site and
D-type site and its adjoining areas, where there are abundant flowering, host,
and nectar-collecting plants and wildflowers, and a minimum number of
butterflies were listed from old and new guest house site and transit
campsites. Among four study years, 2020
had the highest genera and species number followed by the year 2021. From the
present study it can be concluded that despite urbanization, there is a good
diversity of butterflies. Therefore, the implementation of appropriate and
effective conservation methods is of utmost importance in order to protect the
diversity.
Keywords: Conservation
ecology, diversity, ecological indicator, flowering plants, Kamrup District,
Lepidoptera, northeastern India, seasonal variation, species richness,
urbanization.
INTRODUCTION
Butterflies act as an
ecological indicator of environmental variation and are highly sensitive to
disturbances and changes in habitat (Nally & Fleishman 2004). In the field
of conservation ecology, butterflies are considered an umbrella species (Betrus
et al. 2005). It is worth mentioning that butterfly diversity indirectly
indicates plant diversity because both butterfly adults and caterpillars are
highly reliable on specific host plants (Padhye et al. 2006).
In India, 1,379
butterfly species, from six different families, viz., Papilionidae,
Pieridae, Lycaenidae, Riodinidae, Nymphalidae, and Hesperiidae, with 74 endemic
species were observed. Among these, 1,143 species (82.9%) were identified as
Oriental elements, 206 species (14.9%) as Palearctic elements, and 23 species
(1.7%) as Afrotropical elements. Over two-thirds of the species were documented
in the northeastern states of India (Das et al. 2023). Most of the species of
order Lepidoptera indicates meta population which are exclusively phytophagous in
nature (Menken et al. 2010). Northeastern India comprises eight states, viz.,
Arunachal Pradesh, Assam, Manipur, Mizoram, Nagaland, Sikkim, and Tripura are
one the richest biodiversity areas which supports a rich butterfly fauna (Bora
& Meitei 2014). A huge variety of flowering plants, suitable habitats,
topography and climates are ideal for butterfly distribution, diversity and
abundance. Eastern Himalayan part as well as northeastern region of India
comprises 58% of butterflies found in the Indian subcontinent and Myanmar
(Evans 1932). Evans (1932) reported that about 962 species and subspecies of
butterflies belonging to five taxonomic families are found in northeastern
India alone.
Limited research has
been carried out on the butterflies of Assam. A total of 70 species of
butterflies belonging to 45 genera were documented from the Regional Research
Laboratory Campus, Jorhat, Assam (Bhuyan et al. 2005). In various parts of
Guwahati city, a number of studies were conducted to find out the number of
butterfly species. A total of 72 species have been reported from Assam State
Zoo-Cum-Botanical Garden, Guwahati (Ali & Basistha 2000). Saikia et al.
(2015) provided an excellent documentation of about 18 species of butterflies
from Jalukbari and Gauhati university campus, Guwahati. A survey in
Nambor–Doigrung Wildlife Sanctuary, Assam, identified 224 butterfly species
across 137 genera and five families, with Nymphalidae being the most dominant.
A study surveyed butterfly diversity in Dangori Reserve Forest, Upper Assam,
documenting 121 species across six families, with Nymphalidae being the most
dominant. Significant findings include the recording of rare species such as Tanaecia
julii and Lethe chandica, along with endemic species like Arhopala
ganesa and Mycalesis mineus (Boruah & Das 2017). A study
was conducted in Panbari Forest, Kaziranga, upper Assam, that presented a
checklist of 137 skipper butterfly species (Hesperiidae) including species such
as Purple Lancer Salanoemia fuscicornis, Red-vein Lancer Pyroneura
niasana burmana, Pied Flat Celaenorrhinus moreana, and various
species of Choaspes, Potanthus, and Halpe (Gogoi 2013).
Under this
contemplated background, the present study was carried out to identify and
estimate the butterfly diversity in IIT, Guwahati campus.
MATERIALS AND METHODS
Study Area
The present study was
conducted on the IIT Guwahati campus in Assam, renowned as one of the most
beautiful campuses in India. Located at 26.187 N and 91.691 E, the campus lies
on the northern banks of the Brahmaputra River, connected to northern Guwahati’s
Amingaon town, and is approximately 20 km from the city center. Spanning 700 ac
(2.8 km²), the campus features undulating terrain, hillocks, and a variety of
landscapes, including evergreen, semi-evergreen, and deciduous vegetation, as
well as shrubs, grasslands, and wetlands interspersed with lakes.
The campus’s diverse
vegetation, host plants, food plants, and nectar-rich flowers support a rich
variety of reptiles, birds, mammals, insects, and, notably, many vibrant moth
and butterfly species. Study sites included urban habitat areas, hilly
terrains, lakes, wetlands, and specific locations such as the guest house,
administrative block, and serpentile lake. The relatively undisturbed and
tranquil environment of residential and non-residential areas further
contributes to butterfly richness. Detailed descriptions of the selected study
sites are provided in Table 1 and Image 1.
Survey Method
The study was
conducted across various sites at IIT from September 2019 to March 2022 using
the Pollard Walk method. Surveys were performed twice yearly at each site,
between 0900 h and 1700 h on sunny days. Observers walked fixed transects,
recording butterflies within 3–5 m. Unidentified butterflies were caught,
identified using field methods and references, and released (Yasmin et al. 2023). Identification
was primarily done on-site, with photographs used for challenging cases. Data
on date, location, and weather were recorded. The best time for observation and
photography was early mornings, especially after rain showers, when flowering
plants and nectar sources attracted the highest butterfly activity.
The identifications
were done with the help of Evans (1932), and Kehimkar (2008). The WPA, 1972
status of butterflies was obtained from the database available at https://vindhyabachao.org/wildlife_guidelines/schedule_species_insects.pdf
Relative abundance is
calculated by the formula:
Species abundance
Species relative
abundance = ––––––––––––––––– x 100
Total
abundance
RESULTS
During the survey
period from September 2019 to March 2022, a total of 82 species with a total of
1,378 individuals of butterfly belonging to six families and 57 different
genera were recorded from different sites of IIT, Guwahati campus. Checklist of
butterfly species and their abundance in different study sites are shown in
Table 2. The study analyzed the composition of butterfly families over four
years (2019–2022). Nymphalidae emerged as the most dominant family, with the
highest species count and abundance each year, followed by Papilionidae,
Lycaenidae, Pieridae, Hesperiidae, and Riodinidae. Each family displayed
variations in the number of genera, species, and individuals annually, with the
details summarized in Table 3 and Figure 1. Overall, Nymphalidae consistently
led in diversity and population.
The majority of
butterfly species were observed on the old E-type site and the D-type site with
its adjoining areas, which are rich in flowering plants, host plants, and
nectar-collecting wildflowers. In 2019, out of 77 species, 19 were recorded at
the old E-type site, while 14 were found at the D-type site and nearby areas.
In 2020, out of 83 species, both sites recorded 17 species each. Similarly, in
2021 and 2022, out of 79 and 71 species respectively, 20 and 15 species were
found at the old E-type site, while 14 and 16 species were recorded at the
D-type site and its surroundings (Table 4). Figure 2 illustrates butterfly
abundance over four years, showing that 2020 had the highest number of genera
and species, followed by 2021.
Table 5 presents the
relative abundance of butterfly species, while Table 6 and Figure 3 highlight
the relative abundance of different families. The study found that in 2019, Junonia
atlites had the highest relative abundance (3%), whereas Telicota linna
had the lowest (0.09%). In 2020, 2021, and 2022, Papilio polytes
recorded the highest relative abundance at 2.25%, 2.72%, and 3.35%,
respectively. Conversely, Rapala tara, Sarangesa desahara, and Abisara
neophron had the lowest relative abundance (0.15%) in 2020, while Appias
galba (0.18%) and Orsotriaena medus (0.12%) showed the lowest
relative abundance in 2021 and 2022, respectively. Across all four years, the
family Nymphalidae consistently exhibited the highest relative abundance, while
Riodinidae had the lowest (Figure 2).
During this survey,
15 butterfly species with protected status under the Schedule II (Part H with
serial numbers) of The Wild Life (Protection) Amendment Act, 2022, were
recorded. These include Papilio slateri, Graphium sarpedon sarpedon,
Graphium eurypylus, Graphium aristeus anticrates, Cepora nadina nadina, Artipe
eryx, Poritia hewitsoni, Spindasis lohita, Neptis magadha khasiana, Tanaecia
lepidea, Charaxes bernardus, Melanitis zitenius, Ragadia crisilda, Parthenos
sylvia gambrisius and Lethe insana (Table 2).
DISCUSSION
During the survey, a
total of 82 species with about 1,378 individuals of butterfly belonging to six
families (Papilionidae, Pieridae, Lycaenidae, Nymphalidae, Hesperiidae and
Riodinidae) and 57 different genera were recorded in the study area.
Similar studies were
reported by Gogoi et al. (2023) in Soraipung Range of Dehing Patkai
National Park where they recorded a total of 92 butterfly species from five
families, among which 13 species were classified as protected under different
schedules of the Indian Wildlife (Protection) Act, 1972 but according to Wild
Life (Protection) Amendment Act, 2022, nine species among the list of 13
species are now classified as protected under Schedule I and II.
In the present
investigation, a maximum number of butterflies were recorded in the year 2020
(57 genera and 63 species) and 2021 (56 genera and 79 species). This might have
happened because of less human interference, disturbances and environmental
pollution occurred due to COVID-19 pandemic during the year 2020 and 2021.
Comparatively, a lower number of butterflies were documented in the year 2022
(51 genera and 71 species) and 2019 (53 genera and 77 species). Lower number of
butterflies in 2019 may be due to restoration of day-to-day human activities in
these areas.
In the present study,
Nymphalidae family had the highest number and percentage of species of
butterflies in all four years of study period compared with the other families.
The result of the present survey is in close consortium with the findings of
Ali & Basistha (2000). They documented 72 identified species of butterflies
belonging to five families with the highest number of species of the
Nymphalidae family from Assam State Zoo-cum-Botanical Garden, Guwahati, Assam.
Furthermore, the survey of Bohra & Purkayastha (2021) of the urban
landscape, of Guwahati, Assam, India, listed 249 species of butterflies
belonging to six families. The Nymphalidae family was represented as dominant
during the survey period. Adaptation and proper landscape management could be
the reason for the high diversity of the family Nymphalidae. Another reason for
the rich diversity of the family Nymphalidae might be due to their strong
active flying capability and their polyphagous nature which facilitates them to
cover large areas and utilize a variety of host plants (Eswaran & Pramod
2005; Janz 2005; Padhye et al. 2006).
Faunal diversity is
dependent upon the habitat types, food resources and food quality. The
diversity and distribution of butterfly species are also influenced by
sufficient larval and adult plant resources (Ramesh et al. 2010). In the
present study, the highest relative abundances of different species of
butterfly family were found in old E-type and hill top sites. Even though the
old E-type site is an urbanized area and has human interference, the residents
of this area have transformed the environment of the place in such a way that
it has become an attractive and favourable place for butterflies. In agreement
with the result of the present study, different earlier studies have shown that
butterfly diversity in disturbed habitats is more than in undisturbed areas
(Spitzer et al.1993; Hamer et al. 1997). Hill top is the least disturbed area
and the occurrence of sufficient host plants make it more favourable for
butterflies. Junonia atlites was found to have the highest relative
abundance in 2019 and in the years 2020, 2021 and 2022. Papilio polytes showed
highest relative abundance. Both Junonia atlites and Papilio polytes are
common and most frequently observed butterflies and this may happen due to
their adaptation power and the presence of a large number of host plants.
Adult butterflies
generally prefer forest areas with medium altitude and larvae prefer ecotones
with an abundance of food plants with large leaves (Piccini et al. 2022).
Therefore, to conserve this beautiful creation or Nature’s jewels, the suitable
environment for the butterflies should be maintained as well as enhanced.
Therefore, to restore growth of butterfly population, enough plantation should
be carried out in and around the IIT Guwahati campus area. Prevention of human
interventions and disturbances and also deforestation for the purpose of
clearing land for buildings in the hilly arears and lake sides, by the
management, will be a huge step towards the conservation of these amazing
insects.
The findings of this
study suggest that despite ongoing urbanization within the IIT Guwahati campus
in Assam, the area still supports a thriving diversity of butterflies. During the study period, a total of 82
butterfly species, comprising 1,378 individuals from six families and 57
genera, were documented. Among these families, Nymphalidae exhibited the highest
species count and percentage, followed by Papilionidae, Lycaenidae, Pieridae,
Hesperiidae, and Riodinidae, in descending order of abundance (Nymphalidae >
Papilionidae > Lycaenidae > Pieridae > Hesperiidae > Riodinidae).
Table 1. Types
of habitats present in different study sites of IIT campus of Guwahati, Assam.
|
|
Study site |
Name of the study
site |
Habitats |
|
1 |
Site 1 |
D type and its
adjoining area |
Flowering plants,
urban habitat, children park |
|
2 |
Site 2 |
New E type |
Vegetation,
different plants |
|
3 |
Site 3 |
Old E type |
Urban habitat, lots
of plantation, flowering plants, children park |
|
4 |
Site 4 |
F type and its
adjoining area |
Urban habitat, good
management of naturally growing flowering plants |
|
5 |
Site 5 |
Old and new guest
house site |
Urban habitat,
flowering plants, vegetation |
|
6 |
Site 6 |
Manas hostel site |
Vegetation, small
water body, flowering plants |
|
7 |
Site 7 |
Hill top |
Dense vegetation,
different flowering plants and fruit trees |
|
8 |
Site 8 |
Serpentile lake and
its adjoining area |
Grass beds,
vegetation and plants, road side plantation |
|
9 |
Site 9 |
Transit camp site |
Wild flowering
plants, vegetation |
|
10 |
Site 10 |
Admin site and its
adjoining area |
Two lakes,
vegetation |
Table 2.
Checklist of butterfly species and their abundance in different study areas.
|
|
Common name |
Scientific Name |
Year |
Site |
Status |
Conservation status, 2022 |
|||||
|
2019 |
2020 |
2021 |
2022 |
||||||||
|
Family:
Papilionidae |
|||||||||||
|
1 |
Common Mormon |
Papilio polytes |
4+ |
4+ |
4+ |
3+ |
Site 3, site 7,
site 9 |
Very common |
|
||
|
2 |
Great Mormon |
Papilio memnon
agenor |
3+ |
4+ |
4+ |
3+ |
All sites |
Common |
|
||
|
3 |
Common lime |
Papilio demoleus |
3+ |
3+ |
2+ |
2+ |
Site 1 site 2, site
5 |
Very common |
|
||
|
4 |
Common mime |
Papilio clytia
clytia |
3+ |
4+ |
4+ |
3+ |
Site 1, site 3,
site 6, site 10 |
Common |
|
||
|
5 |
Blue-striped Mime |
Papilio slateri |
̶ |
3+ |
3+ |
2+ |
Site 2, site 7 |
Not rare |
Schedule II |
||
|
6 |
Red Helen |
Papilio helenus |
2+ |
3+ |
3+ |
+ |
Site 1, site 8 |
Not rare |
|
||
|
7 |
Yellow Helen |
Papilio nephelus |
4+ |
4+ |
4+ |
3+ |
Site 3, site 4,
site 8, |
Not rare |
|
||
|
8 |
Common Bluebottle |
Graphium sarpedon
sarpedon |
+ |
+ |
2+ |
+ |
Site 6, site 10 |
Common |
Schedule II (Part H; No. 255) |
||
|
9 |
Fivebar Swordtail |
Graphium antiphates
pompilius |
+ |
2+ |
3+ |
3+ |
Site 4, Site5 |
Not rare |
|
||
|
10 |
Fourbar Swordtail |
Graphium agetes
aestes |
+ |
2+ |
̶ |
̶ |
Site 1, Site 7 |
Rare |
|
||
|
11 |
Common Jay |
Graphium doson |
2+ |
4+ |
3+ |
2+ |
Site 1, site 2,
site 3 |
Not rare |
|
||
|
12 |
Great Jay |
Graphium eurypylus
cheronus |
2+ |
3+ |
2+ |
̶ |
Site 1, site 3 |
Not rare |
Schedule II (Part
H; No. 264) |
||
|
13 |
Great Zebra |
Graphium xenocles
xenocles |
2+ |
2+ |
1+ |
1+ |
site 2, site 6 |
Not rare |
|
||
|
14 |
Chain Swordtail |
Graphium aristeus
anticrates |
2+ |
3+ |
2+ |
̶ |
Site 5, site 6 |
Not rare |
Schedule II (Part H; No. 252) |
||
|
15 |
Common Rose |
Atrophaneuraaristolochiae |
+ |
2+ |
+ |
+ |
Site 3, Site 7 |
Rare |
|
||
|
16 |
Common Birdwing |
Troides Helena
cerberus |
2+ |
3+ |
3+ |
2+ |
Site 1, site 3 |
Not rare |
|
||
|
17 |
White Dragontail |
Lamproptera curius
curius |
3+ |
3+ |
2+ |
2+ |
Site 5, site 6 |
Not rare |
|
||
|
18 |
Great Windmill |
Byasa dasarada
dasarada |
2+ |
3+ |
2+ |
2+ |
Site 2, site 3 |
Not rare |
|
||
|
19 |
Common Banded Awl |
Hasora chromus |
3+ |
4+ |
3+ |
2+ |
Site 1, site 7 |
Common |
|
||
|
20 |
Yellow Gorgon |
Meandrusa payeni
evan |
3+ |
3+ |
2+ |
2+ |
Site 3, Site 5 |
Not rare |
|
||
|
Family: Pieridae |
|||||||||||
|
1 |
Common grass yellow |
Eurema hacabe |
3+ |
4+ |
3+ |
2+ |
Site 3, site 7,
site 10 |
Very common |
|
||
|
2 |
Great orangetip |
Hebomoia glaucippe |
3+ |
3+ |
2+ |
2+ |
Site 1, site 3 |
Common |
|
||
|
3 |
Common Emigrant |
Catopsilia Pomona |
3+ |
4+ |
3+ |
3+ |
Site 7, site 9 |
Very common |
|
||
|
4 |
Mottled emigrant |
Catopsilia pyranthe |
2+ |
3+ |
2+ |
2+ |
Site 3, site 7 |
Very common |
|
||
|
5 |
Orange Albatross |
Appias galba |
+ |
+ |
+ |
̶ |
Site 4 |
Rare |
|
||
|
6 |
Striped Albatross |
Appias olferna |
2+ |
2+ |
+ |
2+ |
Site 4 |
Common |
|
||
|
7 |
Spot Puffin |
Appias lalage |
2+ |
+ |
̶ |
+ |
site 3, Site 5 |
Not rare |
|
||
|
8 |
Lesser Gull |
Cepora nadina
nadina |
2+ |
3+ |
3+ |
2+ |
Site 1, Site 3 |
Not rare |
Schedule II (Part
H; No. 286) |
||
|
9 |
Redspot Jezebel |
Delias descombesi |
3+ |
4+ |
4+ |
3+ |
Site 1, site 5,
site 7 |
Common |
|
||
|
Family: Lycaenidae |
|||||||||||
|
1 |
Lesser grass blue |
Zizina otis |
3+ |
4+ |
3+ |
3+ |
Site 3, Site 5,
site 10 |
Not rare |
|
||
|
2 |
Forget me not |
Catochrysops strabo |
3+ |
4+ |
3+ |
2+ |
Site 2, site 6,
site 8 |
Common |
|
||
|
3 |
Zebra blue |
Leptotes plinius |
3+ |
3+ |
2+ |
2+ |
Site 1 site 3 |
Common |
|
||
|
4 |
Yamfly |
Loxura atymnus |
2+ |
2+ |
2+ |
+ |
Site 1, site 3 |
Not rare |
|
||
|
5 |
Assam Flash |
Rapala tara |
̶ |
+ |
2+ |
2+ |
Site 5 |
Rare |
|
||
|
6 |
Green Flash |
Artipe eryx |
2+ |
2+ |
+ |
+ |
Site 7, site 8 |
Not rare |
Schedule II |
||
|
7 |
Dingy Lineblue |
Petrelaea dana |
3+ |
2+ |
2+ |
+ |
Site 4, site 5 |
Common |
|
||
|
8 |
Common Tit |
Hypolycaena erylus
himavantus |
2+ |
+ |
+ |
̶ |
Site 6 |
Common |
|
||
|
9 |
Common Gem |
Poritia hewitsoni |
2+ |
3+ |
2+ |
̶ |
Site1, site 9 |
Not rare |
Schedule II (Part
H, No. 23) |
||
|
10 |
Common Lineblue |
Prosotas nora nora |
4+ |
4+ |
3+ |
2+ |
site 3, Site 7 |
common |
|
||
|
11 |
Long-banded
Silverline |
Spindasis lohita |
3+ |
3+ |
2+ |
3+ |
Site 2, Site 3 |
Not rare |
Schedule II (Part
H, No. 53) |
||
|
Family: Nymphalidae |
|||||||||||
|
1 |
Leopard lacewing |
Cethosia cyane |
3+ |
2+ |
2+ |
+ |
Site 6, site 7 |
Not rare |
|
||
|
2 |
Peacock pansy |
Junonia almana |
+ |
2+ |
+ |
̶ |
Site 1, site 3 |
Not rare |
|
||
|
3 |
Lemon pansy |
Junonia lemonias |
4+ |
3+ |
2+ |
2+ |
Site 5, site 8,
site 10 |
Very common |
|
||
|
4 |
Grey pansy |
Junonia atlites |
4+ |
3+ |
3+ |
3+ |
Site 2, site 10 |
Very common |
|
||
|
5 |
Yellow pansy |
Junonia hiertia |
3+ |
4+ |
3+ |
2+ |
Site 7, site 5 |
Common |
|
||
|
6 |
Chocolate Soldier |
Junonia iphita |
2+ |
3+ |
2+ |
2+ |
Site 3, site 6 |
Common |
|
||
|
7 |
Great eggfly |
Hypolimnasbolina |
3+ |
3+ |
3+ |
2+ |
Site 1, site 3,
site 7 |
Very common |
|
||
|
8 |
Plain Tiger |
Danaus chrysippus |
3+ |
3+ |
2+ |
2+ |
Site 5, site 9 |
Common |
|
||
|
9 |
Common four ring |
Ypthima huebneri |
3+ |
4+ |
4+ |
3+ |
Site 3, site 6 |
Common |
|
||
|
10 |
Common fivering |
Ypthima baldus |
2+ |
2+ |
̶ |
+ |
Site 6 |
Common |
|
||
|
11 |
Common palmfly |
Elymnias
hypermnestra |
̶ |
+ |
+ |
̶ |
Site 5 |
Rare |
|
||
|
12 |
Common Bushbrown |
Mycalesis persius |
4+ |
4+ |
3+ |
3+ |
Site 1, site 10 |
Very common |
|
||
|
13 |
Blue tiger |
Tirumala limniace |
3+ |
3+ |
2+ |
2+ |
Site 3, site 5 |
Common |
|
||
|
14 |
Dark Blue Tiger |
Tirumala
septentrionis |
̶ |
+ |
2+ |
+ |
Site 2, site 4 |
Not rare |
|
||
|
15 |
Medus Brown |
Orsotriaena medus |
2+ |
+ |
+ |
+ |
Site 1, site 7 |
Rare |
|
||
|
16 |
Common Indian crow |
Euploea core |
4+ |
3+ |
2+ |
2+ |
Site 5, Site 9 |
Common |
|
||
|
17 |
Common sailor |
Neptis hylas astola |
3+ |
2+ |
3+ |
2+ |
Site 1, site 3,
Site 10 |
Common |
|
||
|
18 |
Spotted Sailor |
Neptis magadha
khasiana |
+ |
2+ |
+ |
2+ |
Site 3, site 5 |
Rare |
Schedule II (Part
H; No. 216) |
||
|
19 |
Grey count |
Tanaecia lepidea |
2+ |
4+ |
3+ |
3+ |
Site 1, site 4 |
Common |
Schedule II (Part
H; No. 165) |
||
|
20 |
Plain Earl |
Tanaecia jahnu |
+ |
2+ |
+ |
+ |
Site 3, Site 6 |
Rare |
|
||
|
21 |
Colour sergeant
female |
Athyma nefte |
3+ |
2+ |
+ |
2+ |
Site 5, site 6 |
Rare |
|
||
|
22 |
Tawny Rajah |
Charaxes bernardus |
2+ |
4+ |
3+ |
3+ |
Site 7, site 10 |
Common |
Schedule II (Part
H; No. 223) |
||
|
23 |
Tawny coster |
Acraea terpsicore |
2+ |
4+ |
3+ |
2+ |
Site 3, site 5 |
Common |
|
||
|
24 |
Common Evening Brown |
Melanitis leda |
3+ |
2+ |
2+ |
+ |
Site 6, site 10 |
Common |
|
||
|
25 |
Great Evening Brown |
Melanitis zitenius |
+ |
2+ |
2+ |
2+ |
Site 1, site 3 |
Rare |
Schedule II (Part
H; No. 160) |
||
|
26 |
Plain Tiger |
Danaus chrysippus |
3+ |
3+ |
2+ |
+ |
Site 3, Site 8 |
Very common |
|
||
|
27 |
Glassy Tiger |
Parantica aglea |
+ |
2+ |
4+ |
2+ |
Site 3, site 7 |
Common |
|
||
|
28 |
Vagrant |
Vagrans egista |
2+ |
+ |
̶ |
2+ |
Site 6 |
Not rare |
|
||
|
29 |
Common Sergeant |
Athyma perius |
2+ |
3+ |
2+ |
2+ |
Site 8 |
Common |
|
||
|
30 |
Striped Ringlet |
Ragadia crisilda |
̶ |
2+ |
2+ |
2+ |
Site 2, site 6 |
Not rare |
Schedule II |
||
|
31 |
Clipper |
Parthenos sylvia
gambrisius |
2+ |
2+ |
+ |
̶ |
Site 2, Site 5 |
Rare |
Schedule II (Part
H; No. 135) |
||
|
32 |
Common Forester |
Lethe insana |
3+ |
+ |
+ |
- |
Site 1, site 5,
site 7 |
Not rare |
Schedule II (Part
H; No. 141) |
||
|
33 |
Great Nawab |
Polyura eudamippus
eudamippus |
+ |
2 + |
+ |
2+ |
site 1, Site 3 |
Rare |
|
||
|
34 |
Rustic |
Cupha erymanthis
lotis |
2+ |
3+ |
3+ |
2+ |
Site 6 |
Not rare |
|
||
|
Family: Hesperiidae |
|||||||||||
|
1 |
Common small flat |
Sarangesa desahara |
̶ |
+ |
2+ |
+ |
Site 2, site 5 |
rare |
|
||
|
2 |
Common snow flat |
Tagiades japetus
atticus |
4+ |
4+ |
3+ |
2+ |
Site 3, site 10 |
Common |
|
||
|
3 |
Paint Brush Swift |
Baoris farri |
3+ |
3+ |
2+ |
2+ |
Site 1, Site 7 |
common |
|
||
|
4 |
Great Swift |
Pelopidas
assamensis |
+ |
2+ |
+ |
̶ |
Site 3, site 4 |
Not rare |
|
||
|
5 |
Linna Palm Dart |
Telicota linna |
+ |
2+ |
2+ |
̶ |
Site 3, site 7 |
Common |
|
||
|
Family: Riodinidae |
|||||||||||
|
1 |
Punchinello |
Zemeros flegyas
indicus |
3+ |
4+ |
3+ |
2+ |
Site 3, site 10 |
Common |
|
||
|
2 |
Double Banded Judy |
Abisara bifasciata |
3+ |
2+ |
2+ |
2+ |
Site 1, site 3 |
Not rare |
|
||
|
3 |
Tailed Judy |
Abisara neophron |
2+ |
+ |
+ |
- |
Site 2, site 10 |
Not rare |
|
||
N.B. On the basis of abundance, Butterfly species were included under
classes: 4+ (highly abundant, more than
25 sightings), 3+(moderately abundant, 16–25 sighting), 2+(abundant, 6–15
sighting); +(present, 1–5 sighting), -(absent).
Table 3. Number
of genera and species of five families of butterfly.
|
Family |
Year |
|||||||||||||||
|
2019 |
2020 |
2021 |
2022 |
|||||||||||||
|
No. of Genera |
% |
No. of Species |
% |
No. of Genera |
% |
No. of Species |
% |
No. of
Genera |
% |
No. of Species |
% |
No. of genera |
% |
No. of Species |
% |
|
|
Papilionidae |
9 |
16.98 |
21 |
27.27 |
9 |
15.79 |
22 |
26.51 |
9 |
16.07 |
21 |
26.58 |
9 |
17.65 |
19 |
26.76 |
|
Pieridae |
6 |
11.32 |
8 |
10.39 |
6 |
10.53 |
8 |
9.64 |
6 |
10.71 |
7 |
8.86 |
6 |
11.76 |
7 |
9.86 |
|
Lycaenidae |
10 |
18.87 |
10 |
12.99 |
11 |
19.30 |
11 |
13.25 |
11 |
19.64 |
11 |
13.92 |
9 |
17.64 |
9 |
12.68 |
|
Nymphalidae |
22 |
41.51 |
31 |
40.26 |
24 |
42.11 |
34 |
40.96 |
23 |
41.07 |
32 |
40.51 |
22 |
43.14 |
31 |
43.66 |
|
Hesperiidae |
4 |
7.55 |
4 |
5.19 |
5 |
8.77 |
5 |
6.02 |
5 |
8.93 |
5 |
6.33 |
3 |
5.88 |
3 |
4.23 |
|
Riodinidae |
2 |
3.77 |
3 |
3.90 |
2 |
3.51 |
3 |
3.61 |
2 |
3.57 |
3 |
3.80 |
2 |
3.92 |
2 |
2.82 |
|
Total = 53 |
Total = 77 |
Total = 57 |
Total = 81 |
Total = 56 |
Total = 79 |
Total = 51 |
Total = 71 |
|||||||||
Table 4.
Number of butterfly species in different sites of study area.
|
Sites |
Name of the site |
2019 |
2020 |
2021 |
2022 |
|
Site 1 |
D-type and its
adjoining area |
14 |
17 |
14 |
16 |
|
Site 2 |
New E-type |
7 |
5 |
2 |
1 |
|
Site 3 |
Old E-type |
19 |
17 |
20 |
15 |
|
Site 4 |
F type and its
adjoining area |
8 |
12 |
15 |
12 |
|
Site 5 |
Old and new guest
house |
1 |
2 |
3 |
1 |
|
Site 6 |
Manas hostel site |
9 |
11 |
8 |
5 |
|
Site 7 |
Hill top |
11 |
13 |
9 |
7 |
|
Site 8 |
Serpentile lake and
its adjoining area |
4 |
2 |
3 |
5 |
|
Site 9 |
Transit camp site |
2 |
1 |
1 |
3 |
|
Site 10 |
Admin site and its
adjoining area |
2 |
3 |
4 |
6 |
Table 5.
Numerical abundance and relative abundance (RA) of individual butterflies
across the study area.
|
|
Common name |
Family |
2019 |
2020 |
2021 |
2022 |
||||
|
No. |
RA (%) |
No. |
RA (%) |
No. |
RA (%) |
No. |
RA (%) |
|||
|
1 |
Common Mormon |
Papilionidae |
27 |
2.45 |
31 |
2.25 |
30 |
2.72 |
28 |
3.35 |
|
2 |
Great Mormon |
24 |
2.18 |
28 |
2.03 |
26 |
2.36 |
23 |
2.75 |
|
|
3 |
Common lime |
19 |
1.73 |
21 |
1.52 |
15 |
1.36 |
13 |
1.55 |
|
|
4 |
Common mime |
22 |
2 |
29 |
2.1 |
27 |
2.45 |
24 |
2.89 |
|
|
5 |
Blue-striped Mime |
- |
- |
18 |
1.31 |
16 |
1.45 |
13 |
1.55 |
|
|
6 |
Red Helen |
7 |
0.64 |
16 |
1.16 |
17 |
1.54 |
5 |
0.6 |
|
|
7 |
Yellow Helen |
27 |
2.45 |
29 |
2.1 |
28 |
2.54 |
23 |
2.75 |
|
|
8 |
Common Bluebottle |
2 |
0.18 |
5 |
0.36 |
7 |
0.64 |
2 |
0.24 |
|
|
9 |
Fivebar Swordtail |
5 |
0.45 |
9 |
0.65 |
16 |
1.45 |
19 |
2.27 |
|
|
10 |
Fourbar Swordtail |
3 |
0.27 |
7 |
0.51 |
- |
- |
- |
- |
|
|
11 |
Common Jay |
14 |
1.27 |
26 |
1.89 |
23 |
2.09 |
16 |
1.91 |
|
|
12 |
Great Jay |
9 |
0.82 |
18 |
1.31 |
11 |
1 |
- |
- |
|
|
13 |
Great Zebra |
11 |
1 |
15 |
1.09 |
12 |
1.09 |
9 |
1.08 |
|
|
14 |
Chain Swordtail |
7 |
0.64 |
15 |
1.09 |
6 |
0.54 |
- |
- |
|
|
15 |
Common Rose |
3 |
0.27 |
9 |
0.65 |
5 |
0.45 |
3 |
0.36 |
|
|
16 |
Common Birdwing |
12 |
1.09 |
17 |
1.23 |
19 |
1.72 |
15 |
1.8 |
|
|
17 |
White Dragontail |
17 |
1.55 |
20 |
1.45 |
14 |
1.27 |
12 |
1.43 |
|
|
18 |
Great Windmill |
8 |
0.73 |
18 |
1.31 |
10 |
0.91 |
11 |
1.31 |
|
|
19 |
Common Banded Awl |
21 |
1.91 |
27 |
1.96 |
24 |
2.18 |
13 |
1.55 |
|
|
20 |
Yellow Gorgon |
21 |
1.91 |
19 |
1.38 |
13 |
1.18 |
10 |
1.19 |
|
|
21 |
Common grass yellow |
Pieridae |
21 |
1.91 |
29 |
2.1 |
26 |
2.36 |
15 |
1.8 |
|
22 |
Great orangetip |
19 |
1.73 |
21 |
1.52 |
15 |
1.36 |
11 |
1.31 |
|
|
23 |
Common Emigrant |
25 |
2.27 |
27 |
1.96 |
23 |
2.09 |
22 |
2.63 |
|
|
24 |
Mottled emigrant |
13 |
1.18 |
18 |
1.31 |
15 |
1.36 |
14 |
1.67 |
|
|
25 |
Orange Albatross |
8 |
0.73 |
5 |
0.36 |
2 |
0.18 |
- |
- |
|
|
26 |
Striped Albatross |
12 |
1.09 |
13 |
0.94 |
11 |
1 |
14 |
1.67 |
|
|
27 |
Spot Puffin |
6 |
0.55 |
4 |
0.29 |
- |
- |
2 |
0.24 |
|
|
28 |
Lesser Gull |
11 |
1 |
16 |
1.16 |
17 |
1.54 |
12 |
1.43 |
|
|
29 |
Redspot Jezebel |
19 |
1.72 |
26 |
1.89 |
29 |
2.63 |
21 |
2.51 |
|
|
30 |
Lesser grass
blue |
Lycaenidae |
24 |
2.18 |
28 |
2.03 |
24 |
2.18 |
20 |
2.39 |
|
31 |
Forget me not |
25 |
2.27 |
31 |
2.25 |
21 |
1.91 |
14 |
1.67 |
|
|
32 |
Zebra blue |
17 |
1.55 |
22 |
1.6 |
11 |
1 |
9 |
1.08 |
|
|
33 |
Yamfly |
5 |
0.45 |
7 |
0.51 |
4 |
0.36 |
3 |
0.36 |
|
|
34 |
Assam Flash |
- |
- |
2 |
0.15 |
7 |
0.64 |
5 |
0.6 |
|
|
35 |
Green Flash |
8 |
0.73 |
5 |
0.36 |
3 |
0.27 |
2 |
0.24 |
|
|
36 |
Dingy Lineblue |
17 |
1.55 |
13 |
0.94 |
12 |
1.09 |
5 |
0.6 |
|
|
37 |
Common Tit |
7 |
0.64 |
4 |
0.29 |
4 |
0.36 |
3 |
0.36 |
|
|
38 |
Common Gem |
|
8 |
0.73 |
16 |
1.16 |
6 |
0.54 |
- |
- |
|
39 |
Common Lineblue |
27 |
2.45 |
30 |
2.18 |
25 |
2.27 |
14 |
1.67 |
|
|
40 |
Long-banded
Silverline |
17 |
1.55 |
23 |
1.67 |
13 |
1.18 |
27 |
3.23 |
|
|
41 |
Leopard lacewing |
Nymphalidae |
21 |
1.91 |
15 |
1.09 |
10 |
0.91 |
5 |
0.6 |
|
42 |
Peacock pancy |
3 |
0.27 |
8 |
0.58 |
4 |
0.36 |
- |
- |
|
|
43 |
Lemon pancy |
29 |
2.64 |
22 |
1.6 |
14 |
1.27 |
9 |
1.08 |
|
|
44 |
Grey pancy |
33 |
3 |
25 |
1.81 |
25 |
2.27 |
24 |
2.87 |
|
|
45 |
Yellow pansy |
24 |
2.18 |
30 |
2.18 |
21 |
1.91 |
15 |
1.79 |
|
|
46 |
Chocolate Soldier |
8 |
0.73 |
19 |
1.38 |
16 |
1.45 |
14 |
1.67 |
|
|
47 |
Great eggfly |
16 |
1.45 |
22 |
1.6 |
19 |
1.72 |
14 |
1.67 |
|
|
48 |
Danaid eggfly |
16 |
1.45 |
18 |
1.31 |
11 |
1 |
8 |
0.96 |
|
|
49 |
Common fouring |
23 |
2.09 |
28 |
2.03 |
28 |
2.54 |
25 |
2.99 |
|
|
50 |
Common fivering |
7 |
0.64 |
6 |
0.44 |
- |
- |
2 |
0.24 |
|
|
51 |
Blue striped
palmfly |
- |
- |
5 |
0.36 |
3 |
0.27 |
- |
- |
|
|
52 |
Common Bushbrown |
27 |
2.45 |
31 |
2.25 |
29 |
2.63 |
23 |
2.75 |
|
|
53 |
Blue tiger |
20 |
1.82 |
24 |
1.74 |
13 |
1.18 |
14 |
1.67 |
|
|
54 |
Dark Blue Tiger |
- |
- |
3 |
0.22 |
8 |
0.73 |
5 |
0.6 |
|
|
55 |
Pointed Palmfly |
9 |
0.82 |
7 |
0.51 |
3 |
0.27 |
1 |
0.12 |
|
|
56 |
Common Indian crow |
27 |
2.45 |
21 |
1.52 |
13 |
1.18 |
14 |
1.67 |
|
|
57 |
Common sailor |
20 |
1.82 |
14 |
1.02 |
19 |
1.72 |
12 |
1.43 |
|
|
58 |
Spotted Sailor |
5 |
0.45 |
11 |
0.79 |
4 |
0.36 |
9 |
1.08 |
|
|
59 |
Grey count |
13 |
1.18 |
27 |
1.96 |
22 |
2 |
19 |
2.27 |
|
|
60 |
Plain Earl |
2 |
0.18 |
6 |
0.44 |
3 |
0.27 |
3 |
0.36 |
|
|
61 |
Perak Lascar |
23 |
2.09 |
12 |
0.87 |
5 |
0.45 |
11 |
1.31 |
|
|
62 |
Tawny Rajah |
13 |
1.18 |
28 |
2.03 |
20 |
1.81 |
14 |
1.67 |
|
|
63 |
Tawny coster |
15 |
1.36 |
29 |
2.10 |
25 |
2.27 |
15 |
1.79 |
|
|
64 |
Common EveningBrown
|
21 |
1.91 |
9 |
0.65 |
10 |
0.91 |
4 |
0.48 |
|
|
65 |
Great EveningBrown |
2 |
0.18 |
11 |
0.79 |
8 |
0.73 |
6 |
0.72 |
|
|
66 |
Plain Tiger |
18 |
1.64 |
17 |
1.23 |
9 |
0.82 |
5 |
0.6 |
|
|
67 |
Glassy Tiger |
5 |
0.45 |
14 |
1.02 |
27 |
2.45 |
13 |
1.55 |
|
|
68 |
Vagrant |
7 |
0.64 |
3 |
0.22 |
- |
- |
9 |
1.08 |
|
|
69 |
Common Sergeant |
13 |
1.18 |
24 |
1.74 |
14 |
1.27 |
12 |
1.43 |
|
|
70 |
Striped
Ringlet |
- |
- |
6 |
0.44 |
10 |
0.91 |
8 |
0.96 |
|
|
71 |
Clipper |
11 |
1 |
8 |
0.58 |
4 |
0.36 |
- |
- |
|
|
72 |
Common Forester |
14 |
1.27 |
23 |
1.67 |
9 |
0.82 |
4 |
0.48 |
|
|
73 |
Great Nawab |
2 |
0.18 |
8 |
0.58 |
5 |
0.45 |
9 |
1.08 |
|
|
74 |
Rustic |
12 |
1.09 |
21 |
1.52 |
19 |
1.72 |
13 |
1.55 |
|
|
75 |
Common small flat |
Hesperiidae |
- |
- |
2 |
0.15 |
7 |
0.64 |
4 |
0.48 |
|
76 |
Common snow flat |
26 |
2.36 |
27 |
1.96 |
23 |
2.09 |
15 |
1.79 |
|
|
77 |
Paint Brush Swift |
20 |
1.82 |
18 |
1.31 |
8 |
0.73 |
7 |
0.84 |
|
|
78 |
Great Swift |
3 |
0.27 |
9 |
0.65 |
4 |
0.36 |
- |
- |
|
|
79 |
Linna Palm Dart |
|
1 |
0.09 |
6 |
0.44 |
3 |
0.27 |
- |
- |
|
80 |
Punchinello |
Riodinidae |
21 |
1.91 |
30 |
2.18 |
22 |
2 |
19 |
2.27 |
|
81 |
Plum Judy |
17 |
1.55 |
12 |
0.87 |
13 |
1.18 |
9 |
1.08 |
|
|
82 |
Tailed Judy |
8 |
0.73 |
2 |
0.15 |
3 |
0.27 |
- |
- |
|
For
figures & images - - click here for full PDF
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