Butterflies
(Lepidoptera: Papilionoidea and Hesperoidea) and other protected fauna of Jones
Estate, a dying watershed in the Kumaon Himalaya, Uttarakhand, India
Peter Smetacek
Butterfly Research Centre, The Retreat, Jones Estate, Bhimtal, Uttarakhand
263136, India
Email: petersmetacek@rediffmail.com
Date of publication (online): 26 August 2012
Date of publication (print): 26 August 2012
ISSN 0974-7907 (online) | 0974-7893 (print)
Editor: Rudi Mattoni
Manuscript details:
Ms # o3020
Received 25 November 2011
Final received 02 February 2012
Finally accepted 15 July 2012
Citation: Smetacek, P. (2012). Butterflies
(Lepidoptera: Papilionoidea and Hesperoidea) and other protected fauna of Jones
Estate, a dying watershed in the Kumaon Himalaya, Uttarakhand, India. Journal of Threatened Taxa 4(9): 2857–2874.
Copyright: © Peter Smetacek 2012. Creative Commons Attribution 3.0 Unported License. JoTT
allows unrestricted use of this article in any medium for non-profit purposes,
reproduction and distribution by providing adequate credit to the authors and
the source of publication.
Author Details: Peter Smetacek is an authority on Indian
Lepidoptera and has pioneered the use of insect communities as bio-indicators
of climatic change and ground water.
Acknowledgements: I am grateful to my late
father, Fred Smetacek Sr.; to the Times Fellowship
Council, New Delhi, for a Fellowship to study Indian rivers in 1992 and to the
Rufford Small Grant Foundation, U.K., for funding the work on Lepidoptera and
Himalayan forest ecosystems between 2006 and the present
study via a series of grants. I am indebted to Rudi Mattoni, Argentina for
encouragement to write this and valuable suggestions on an earlier draft and
drawing my attention to the format developed by him and used in Table 1 and to
Zdenek Faltynek Fric, Ceske Budejovice in the Czech Republic, for valuable taxonomic
comments on Table 1. Also, I am indebted to the anonymous referees whose
recommendations considerably improved the paper and to my children, Kanika and
Pius, who spent many hours sorting through note books, loose leaf lists, books
and specimens to compile the enormous amount of data that went into the making
of Table 1.
Abstract:Two hundred and forty three species of butterflies recorded from Jones
Estate, Uttarakhand between 1951 and 2010 are reported. The ongoing rapid
urbanization of Jones Estate micro-watershed will destroy the habitat of 49
species of wildlife protected under Indian law, as well as several species of
narrow endemic moths and butterflies. The only known Indian habitat for the butterfly Lister’s Hairstreak Pamela dudgeoni will be
destroyed. The effect on the water
flow of both the Bhimtal and Sattal lake systems will clearly be adverse, as is
evident from the drying up of Kua Tal and the reduced flow of perennial water
springs during the dry season on the Estate. The undoubtedly negative effect of
urbanization on these valuable fresh water resources will be irreversible in
the long term. The trend can be
reversed by extending protection to Jones Estate by re-declaring it a Green Belt
of Bhimtal and by banning construction in the catchment area of Bhimtal lake,
as has been done in Nainital and Mussoorie, both in Uttarakhand.
Keywords: Bhimtal,
drinking water, drying lakes, freshwater resources, Green Belt, Indian Wildlife
(Protection) Act 1972, Pamela
dudgeoni.
For figures, images, tables --
click here
INTRODUCTION
Jones Estate (“June State” on Revenue
Department records) is a forested microwatershed in Nainital district,
Uttarakhand (29021’17”N & 79032’34.27”E),
separating the Bhimtal and Sattal lake systems (Image 1). In the Himalaya, it is a unique
geographical feature, being the only forested watershed separating two lake
systems comprising a total of eight perennial and seasonal lakes. Bhimtal (tal = lake in Hindi) lies at
the southeastern end while the Sattal lies along the northern half of the
western face of the Estate. Comprising roughly 4.8 sq.km (1200 acres) of private forest in 1951, the
forest area of Jones Estate has been reduced by roughly 30% due to cultivation
and habitation over the years.
The lowest point is 1200m at the
conjoined Ram and Sita lakes of the Sattal (seven lakes) system, while the
highest point is Thala at 1731m. The range runs northwest to southeast for a distance of roughly 3km from
Dhupchaura pass on the northwest to Tallital market and Bohrakun Village on its
southeastern and southern faces
respectively (Image 1). To the east
lies the Bhimtal lake system comprising of three lakes—Nal-Damayantital,
Kuatal and Bhimtal. To the west
lies the Sattal lake system comprising of Pannatal (=Garurtal) (Image 2); Ramtal and Sitatal, Lakshmantal, Sukhatal, Sariyatal and
Lokhamtal.
Jones Estate lies in the outermost range
of the Himalayan foothills and receives heavy rainfall. Although Osmaston (1927) gives a range
of 2000–3000 mm of rainfall for this area, actual precipitation is rather
less nowadays, averaging 1443mm for the five-year period from 2005 to 2009
(Anonymous 2010).
The forest consists of three plant
associations, namely sub-tropical broadleaf with Himalayan Oak Quercus
leucotrichophora as a nodal species; Chir Pine Pinus
roxburghii forest and elements of miscellaneous deciduous forest. In addition, there is a patch of
naturalized Himalayan Cypress Cupressus torulosa several acres in
extent.
HISTORY
The Bhimtal Valley has been inhabited and
cultivated for over a millennium and Atkinson (1882) noted that it was one of
the largest single sheets of cultivation in
the Kumaon Himalaya.
Jones Estate watershed and the Sattal
Valley were not inhabited during past centuries, although some small patches of
cultivation were attempted by share-croppers and
itinerant families until 1952. The
major part of the Estate has always been forested. It came into existence in
1867 as a fee-simple estate, with the main aim
of developing it for the production of green tea for the Tibetan market. Since then, it has remained in private
hands.
On 17 January 2001 the then Minister for
Environment and Forests, Mr. Kandari, stated in the Uttarakhand State Assembly
that all concerned government departments, including the Public Works
Department, Forest Department, Pollution Control Department, etc. in their
reports on the possibility of construction of buildings on the Jones Estate
watershed, had stressed that any such move would result in disastrous
consequences for Bhimtal and for the water storage capability of the Sattal
lakes (Special Correspondent 2001).
In 1954, a Forest Working Plan was passed
for Jones Estate by the Forest Department. In the Land Record Settlement known as the Bandobast in 1957, land use
of the greater part of Jones Estate was recorded as “forest”. Under the provisions of the Forest
Conservation Act 1980, no land use change is permitted on such land without the
permission of the Central Government.
Despite this, numerous houses and resorts
are being constructed and there is no doubt that
the eventual urbanization of the Estate is well under way, to the detriment of
the Bhimtal and Sattal lake systems and the wildlife inhabiting the Estate at
present. Therefore, the present
paper documents the butterflies recorded on the Estate (Table 1) so as to get a
better idea of what is being lost and the eventual consequences of urbanization
of the Jones Estate watershed.
The present paper also documents the
butterflies and vertebrates afforded
protection under the Schedules of the Indian Wildlife (Protection) Act 1972
(Anonymous 2006) that have been recorded on the Estate (Table 2). In total, these constitute 49 species,
11 on Schedule 1 and 39 on Schedule 2 (Hypolimnas misippus Linnaeus
figures on both Schedules and is counted only once).
MATERIAL AND METHODS
The butterflies of the Estate have been
studied since 1951. Some original
specimens still exist, but the major resource from this era is in the form of
notes maintained by my father, the late Fred Smetacek Sr. In the course of studying local
butterflies, two butterfly subspecies new to science were discovered on the
Estate, namely Neptis miah varshneyi Smetacek and Neptis clinia
praedicta Smetacek (both Nymphalidae) (Smetacek 2002; 2011b). Besides, several butterflies
previously unrecorded from the Western Himalaya have been reported (Smetacek
2010). Moths have been studied
since 1972. Several species new to
science have been described from the Estate (Smetacek 2002; 2005; 2010a). Besides, the population of hawkmoths
(Lepidoptera: Sphingidae) on the Estate provided material for pioneering work
in using members of an insect community as bio-indicators to predict and track
climate change (Smetacek 1994; 2004).
The sightings of mammals and birds
included in Annexure 2 were compiled mostly during the 1980s, when much time
was spent patrolling the forest. They are all based on actual sightings by the author. The most recent sighting of a mammal protected
under the provisions of the Wildlife (Protection) Act 1972 and listed as Near
Threatened on the IUCN Red Data List is of a Himalayan Serow (Capricornis
sumatraensis thar) which was sighted and photographed outside the Butterfly
Research Centre, Jones Estate at 10am on 07 November 2011.
The use of a lepidopteran community as an
indicator of forest health and consequently the health of the ecosystem
including sub-surface water resources has
been explored on the Estate by the author for the past 30 years (Smetacek
1993–2010). The format of
Image 1, which enables a great deal of information to be presented concisely,
has been taken with kind permission from Mattoni (1990) and Mattoni &
Vannucci (2008), for which I am grateful.
The information recorded in Table 1
regarding voltinism is based on specimens recorded or bred, while the information regarding diapause is based on breeding experiments
by Fred Smetacek Sr. and the author.
WATERSHED FUNCTIONS
Both the Bhimtal and Sattal lake systems
depend largely on subsurface water from this watershed to sustain them. This fact was acknowledged by Atkinson
(1882) in the words, “The hills on the western side of the [Bhimtal] lake [i.e.
present day Jones Estate] are considerably higher than on the east, and are of
such formation that it is highly improbable that the lake can ever dry up.” Of course, back then,it was improbable that Mr. Atkinson could even conceive of the present threat
of urbanization facing Bhimtal in
general and Jones Estate in particular (Image 3).
Today, one lake of the system, Kuatal,
has actually dried up during the last decade. With regard to Kuatal, the only
thing that has changed during the past 30 years is that the Himalayan Oak
forest in its catchment area in Jones Estate has been severely degraded (Image
4). In the wake of nearly
unprecedented rain during July and September 2010, Kuatal filled twice for the
first time in over 60 years. A
photograph from around 1890 (Image 5) shows the area as it apparently used to
be and Image 6 shows Kuatal when it was full in 2010. The loss of Kuatal is
not recognized by any level of Government. Government maps used by the
Lakes Development Authority and other Governmental departments do not even have
a lake marked at the spot.
The only over-ground feeder for any of
the lakes, is a small stream that rises on the Aru
Plain north of Bhimtal and enters the northern end of Bhimtal after a course of
roughly 3km, of which nearly 1km is underground. This stream entirely dried up for two
months between May 2012 and July 2012, for the first time in history. This is
an extremely worrisome development since the prospect of Bhimtal lake becoming seasonal in the coming years is a very real
probability. For the first time,
too, water supply from Bhimtal Lake to Haldwani City was extended from
15–30 June 2012 by the State
Irrigation Department due to the lack of alternate water resources to supply
the city. In addition to the
subsurface springs feeding the lakes, there are two over-ground water springs
on the eastern face of the watershed and three on its western face. In
addition, there are several sub surface springs on the lake
beds of both lake systems. Of the over-ground springs, both eastern face springs used to be
perennial but one is now almost seasonal, with its flow reduced to a thin
trickle during the dry season. Similarly, all of the three western-face springs, which used to be
perennial, are seasonal today, due, presumably, to depleted Oak forest in the
catchment areas of these springs. It is clear that as long as the Oak forest
was in good condition, the springs were perennial. Changes to the Oak forest ecosystem in
the form of the large scale drying up of trees due to forest fires, destruction
of the canopy due to lopping for fodder, cutting of trees, etc were reflected
in reduced flow during the dry season and increased flow, sometimes even
forming new outlets, during the wet season.
DISCUSSION
The Jones Estate watershed separates two
lake systems which together constitute five perennial
lakes and several seasonal ones. These lakes are an inestimably valuable resource of drinking quality
water for future generations. However, with the urbanization of the Jones Estate watershed, their
future looks uncertain, especially the Sattal lake system, which has no
over-ground tributaries and a very small catchment area.
Of the 400+ species of butterflies known
from Uttarakhand (Mackinnon & de Nicéville 1899; Hannyngton 1910, 1915;
Smetacek 1987, 1992, 1993a, 1995, 1999, 2001, 2002, in press b, unpub. data;
Singh 2003, 2005, 2005a, 2006, 2007), 243 species of butterflies (Annexure 1)
and approximately 550 species of macro-moths (Smetacek 2008, 2009) have been
recorded from the Jones Estate watershed. This may be compared with the entire Indian Peninsula, which is home to
330 species of butterflies (Gaonkar 1996)
and an undetermined number of moth species.
Forty nine species of wildlife included in the
Schedules of the Indian Wildlife Protection Act, 1972 have been recorded on the
watershed (Table 2), yet even this impressive figure is not enough to prevent
the ongoing destruction of this forest ecosystem.
Of special interest is the record of
Lister’s Hairstreak Pamela dudgeoni de Nicéville (=Listeria dudgeoni)
(Lepidoptera: Lycaenidae) in March 1974 in Jones Estate. This butterfly was known from Sikkim
(Type locality: specimen described in 1894) and Mussoorie (Evans 1932). Wynter-Blyth (1957) noted a specimen
from Siuni camping ground near Ranikhet in Kumaon. The single specimen was recorded in 1909
and is currently in the collection of the Natural History Museum, London. It has not been recorded from Sikkim
since the type was reported and was evidently recorded from Mussoorie between
1899 and 1932, since Mackinnon & de Nicéville (1899) do not report it. The male specimen on
whose basis Evans (1932) reported it from 1829m (6000ft)
elevation in Mussooorie is currently in the collection of the Forest
Research Institute, Dehradun. Sidhu
(2011) has reported the substantial loss of habitat for the Lycaenidae in
Mussoorie. It is almost certain
that Pamela dudgeoni will not be found in that area now. This leaves Jones Estate as the only
known habitat where this extremely rare butterfly has been recorded
recently. The genus is monobasic
and was believed to be confined to the Himalaya
(Varshney 1997; 2010) although Inayoshi (1996–2011) has reported it from
Thailand. It is not improbable that the ongoing destruction of the insect’s
habitat in Jones Estate will result in the global extinction of the species.
Besides this, Jones Estate is one of
three known localities for Euaspa ziha de Nicéville (Lepidoptera:
Lycaenidae) (P. Smetacek unpub. data); one of two known localities for Neptis
miah varshneyi Smetacek; one of two known localities for Comostola hauensteiniSmetacek (Lepidoptera: Geometridae) (Smetacek 2002). It supports one of the two known populations
of Hestina persimilis Westwood in Uttarakhand (Schedule 2, Indian
Wildlife (Protection) Act 1972) (P. Smetacek unpub. data).
Populations of most butterfly species
listed in Tables 1 and 2 are now concentrated around the Butterfly Research
Centre, due to the destruction of habitat in other parts of the Estate, prior
to the process of building houses. However, the area where these butterfly and moth species are now
confined is very small and the possibility that they become locally extripated
from the Estate in the near future is a very real probability.
The only way to have prevented the
ongoing destruction and to stabilize the Bhimtal and Sattal lake systems as a
drinking water resource for future generations would be, as earlier, to ban the
construction of houses, resorts and cottages on the Estate. However, lacking Governmental will to do
so, the future outlook for this watershed is bleak from the point of view of
insect communities, water resources, the existing forest ecosystems and
especially for the residents of the areas downstream, who might have relied
upon the Bhimtal and Sattal lakes for a dependable supply of drinking quality
water.
RECOMMENDATIONS
If competent decision making bodies make
the following decisions and ensure their implementation, the gravity of the
situation outlined above can be reduced or reversed in the long term:
1. Re-declare Jones Estate a Green Belt,
with the area extended to include the area outlined in Fig. 1.
2. Establish a competent Central
Government committee to look into the implementation of existing laws,
especially the Forest Conservation Act, 1980 and the bylaws of the Lakes
Development Authority, in Jones Estate, and break down illegal constructions to
restore the status quo ante.
3. Ban the keeping of cattle and goats
within the jurisdiction of the Bhimtal Town Area Committee in the same way this
is banned within the Nainital Municipal Area.
Extrapolating the results of the above
findings to the catchment area of Bhimtal and Sattal lake systems is not
difficult, since the areas are contiguous and within the same altitudinal
limits.
If the Bhimtal and Sattal lakes systems
are to be stabilized in the long term, then the following points may be taken
into consideration:
1. An immediate ban be put into effect on
construction of roads, houses and commercial establishments within the
catchment areas of Bhimtal and Sattal lakes, i.e. extending from the Catholic
Church in Bhowali, east along the crest line to Ghorakhal, thence along the
ridges to the hamlet of Binayak, south along the crest of Karkot and thence
west to Bohrakoon hamlet, then further west to Suriya Gaon and northwards along
the crest of Hidamba (Hirrup) to the crest of Hatchhina and thence again following
the line of crests to the Catholic Church in Bhowali. The same provisions as applied in
Nainital or Mussoorie Municipalities regarding construction bemade applicable within this area.
2. Within this area, so as not to
alienate residents, a provision be made to permit residents for at least two
generations (i.e. 40 years) to construct additional housing for extended
families or build commercial establishments for a source of family income. The verification for this may be made
sensible but strict so that lower officials do not consider the possibility of
issuing fake reports or certificates, as the case may be.
3. A ban on cattle and goat breeding be implemented within the jurisdiction of the Bhimtal Town
Area Committee, on the same lines as that in force within Nainital Cantonment
and Municipality.
4. A ban on free range grazing by cattle
within the catchment area, to be enforced by concerned agencies, i.e. Forest
Department and Van Panchayats.
5. A wood depot beset up in Bhimtal so that residents are able to purchase fuel legally, as in
other small towns throughout Kumaon.
6. If necessary, one or more fodder
depot(s) be set up in the higher reaches of the catchment area, eg. in Farsoli, to cater to the needs of villages in that area
and reduce dependence on lopping forest trees.
All these recommendations are well within
the scope of the Government, require no extra funds
and will have to be implemented in due course in the Bhimtal Valley in any
case. If it is done now, then the
action will come into force before the damage is irreversible, unlike Nainital
and Mussoorie, where the bans came into effect after the damage had been done.
REFERENCES
Anonymous (2006). The Wildlife
(Protection) Act 1972. Natraj Publishers, Dehra
Dun, 235pp.
Anonymous (2010). District rainfall
for last 5 years. District: Nainital. Hydromet Division. India Meteorological
Department.http://www.imd.gov.in/section/hydro/
distrainfall/webrain/uttarakhand/nainital.txt. Accessed on 21 July 2010.
Atkinson, E.T. (1882). The Himalayan
Districts of the North West Provinces of India. Vol. 2,
Chapter 2. Zoology (Invertebrata).GovernmentPress, Allahabad, 87–266pp.
Evans, W.H. (1932). The
Identification of Indian Butterflies. 2nd Edition. Bombay Natural History Society, Bombay, 10+454pp+32pls.
Gaonkar, H. (1996). Butterflies
of the Western Ghats, India including Sri Lanka. A Biodiversity Assessment of A Threatened Mountain System. Centre for Ecological Sciences,
Indian Institute of Science, Bangalore; Zoological Museum, Copenhagen and the
Natural History Museum, London, 89pp.
Hannyngton, F. (1910). The
butterflies of Kumaon. Journal of the Bombay Natural History Society20: 130–142; 361–372.
Hannyngton, F. (1915). Kumaon Butterflies. Journal of the
Bombay Natural History Society 24(1): 197.
Inayoshi, Y. (1996–2011). A Check List of Butterflies in
Indo-China, page on Family Lycaenidaehttp://yutaka.it-n.jp/lyc4/81565001.html (accessed on 09 February 2012).
Mackinnon,
P.W. & L. de Nicéville (1899). List
of butterflies of Mussoorie in the Western Himalayas and neighbouring regions. Journal
of the Bombay Natural History Society 11:
205–221, 368–389, 585–605.
Mattoni, R. (1990). Butterflies of
Greater Los Angeles. Lepidoptera Research
Foundation, Beverly Hills, 1–20pp.
Mattoni, R. & N.
Vannucci (2008). Garden
Butterflies of Buenos Aires. Lepidoptera Research
Foundation, Beverly Hills, 1–24pp.
Osmaston, A.E. (1927). A Forest Flora for
Kumaon. Government Press, Allahabad, 34+605pp.
Robinson, G.S., P.R. Ackery, I.J.
Kitching, G.W. Beccaloni & L. Hernandez (2001). Hostplants of The Moth
and Butterfly Caterpillars of The Oriental Region. The Natural History Museum, London and Southdene Sdn. Bht., Kuala
Lumpur, 744pp.
Sidhu, A.K. (2011). Changing biodiversity scenario in the
Himalayan ecosystem: Mussoorie, Uttarakhand, India as revealed by the study of
blue butterflies (Lycaenidae). Journal of Threatened Taxa3(2): 1559–1563.
Singh, A.P. (2003). Distribution range
extension of bush hopper butterfly, Ampittia dioscorides Fabricius
(Lepidoptera: Hesperidae) into the lower western Himalayas. Indian
Forester 129(8): 1046–1048.
Singh, A.P. (2005). Initial colonization of
Red Pierrot butterfly, Talicada nyseus nyseus Guerin (Lycaenidae) in the
lower western Himalayas: an indicator of the changing environment.
Current Science 89: 41–42.
Singh, A.P. (2005a). Recent records on the
distribution, seasonality and occurrence of Redspot
butterfly, Zesius chrysomallus Hübner from the lower western Himalayas. Journal of the Bombay Natural History Society 102(2):
238–239.
Singh, A.P. (2006). Range extension of Brown Gorgon
butterfly, Meandrusa gyas gyas Westwood
into Kedarnath Musk Deer Reserve, Western Himalayas: A lesser
known species from north-east India. Indian Forester 132(12a):
187–189.
Singh, A.P. (2007). A new
butterfly species of the genus Ypthima Hübner (Nymphalidae: Satyrinae)
from Garhwal Himalaya, India. Journal of the Bombay
Natural History
Society 104(2): 191–194.
Smetacek, P. (1987). A new type of mimicry
in butterflies. Journal of the Bombay Natural History Society:
83: 471.
Smetacek, P. (1992). Record of Plebejus
eversmanni (Staudinger) (Lepidoptera: Lycaenidae) from India. Journal of the
Bombay Natural History Society 89: 385–386.
Smetacek, P. (1993). Towards
the Re-habilitation of Indian rivers. Report submitted to Times
Fellowship Council, New Delhi, 53pp.
Smetacek, P. (1993a). Neptis carticaMoore (Lepidoptera: Nymphalidae) in the U.P. Himalaya. Journal
of the Bombay Natural History Society 90: 527–528.
Smetacek, P. (1994). An annotated list of the
Hawkmoths (Lepidoptera: Sphingidae) of Kumaon, North India: a probable case of
faunal drift. Records of the Zoological Survey of India. Occasional Paper 156: 1–55.
Smetacek, P. (1995). A new
altitudinal and range record for the Copper Flash Butterfly Rapala pheretimusHewitson (Lepidoptera: Lycaenidae). Journal of
the Bombay Natural History Society 92: 127–128.
Smetacek, P. (1999). The
distribution and ecology of Polyura agraria Swinhoe (Lepidoptera:
Nymphalidae) in India. Journal of the Bombay
Natural History Society 96: 487–488.
Smetacek, P. (2001). Resolution
of the controversial western limit of the range of Delias acalis Godart
(Lepidoptera: Pieridae). Journal of the Bombay
Natural History Society 98: 298–300.
Smetacek, P. (2002). The genus PontiaFabricius (Lepidoptera: Pieridae) in the Kumaon Himalaya. Journal of the Bombay Natural History Society 99:
224–231.
Smetacek, P. (2004). Descriptions
of new Lepidoptera from the Kumaon Himalaya. Journal
of the Bombay Natural History Society 101: 269–276.
Smetacek, P. (2004). On the
occurrence of Marumba cristata Butler (Lepidoptera: Sphingidae) in
Shimla, Himachal Pradesh, India. Journal of the
Bombay Natural History Society 101: 171–172.
Smetacek, P. (2005). The
Epipleminae (Lepidoptera: Uraniidae) of the Kumaon Himalaya. Journal of the Bombay Natural History Society 102:
186–194.
Smetacek, P. (2008). Moths recorded from different elevations
in Nainital district, Kumaon Himalaya, India. Bionotes10(1): 5–15.
Smetacek, P. (2009). Additions and corrections
to the list of moths recorded from different elevations in Nainital district,
Kumaon Himalaya, India. Bionotes 11(4): 117–118.
Smetacek, P. (2010a). A new species of CeryxWallengren (Lepidoptera: Arctiidae) from the Kumaon Himalaya. Journal of Threatened Taxa 2(5): 894–895.
Smetacek, P. (2010b). Climate Change and
Himalayan Lepidoptera. Abstract Book. International Workshop on Mountain
Biodiversity and Impacts of Climate Change with special reference to Himalayan
Biodiversity Hotspot. G.B. Pant Institute of Himalayan
Environment and Development, Kosi-Katarma, Almora, 130–134pp.
Smetacek, P. (2011a). Four new Lycaenid
records from the Kumaon Himalaya, India. Journal of
Threatened Taxa 3(2): 1555–1558.
Smetacek, P. (2011b). A
review of West Himalayan Neptini (Lepidoptera: Nympalidae). Journal of the Lepidopterists’ Society 65(3): 153–161.
Special
Correspondent (2001). Permission for construction in
Bhimtal’s Green Belt a mistake: Government accepts onus in State Assembly.(in Hindi) Amar Ujala newspaper for January 18, 2001 Dehra
Dun Edition, page 1 lead story, continued on page 2. (Bhimtal ki harit patti kshetra mein nirman ki anumati galat: Sarkar nay
Vidhan Sabha mein sweekar kiya).
Varshney, R.K. (1997). Index Rhopalocera Indica Part III. Genera of Butterflies from India and neighbouring Countries
[Lepidoptera: Lycaenidae]. Oriental Insects 31:
83–138.
Varshney, R.K. (2010). Genera of Indian
Butterflies. Nature Books India, New Delhi, 186pp.
Wynter-Blyth, M.A. (1957). Butterflies of the
Indian Region. Bombay Natural History Society, Bombay, 20+523pp.
+72pls.