Description
of three new species of the genus Allochthonius Chamberlin,
1929 (Pseudoscorpiones: Pseudotyrannochthoniidae) from China
Junfang Hu 1 &
Feng Zhang 2
1,2 College of Life Sciences, Hebei
University, Baoding, Hebei 071002, China
Email: 1 jfanghu@gmail.com, 2 dudu06042001@163.com
(corresponding author)
Date of publication (online):26 November 2011
Date of publication (print): 26 November 2011
ISSN 0974-7907 (online) | 0974-7893 (print)
Editor: Mark S. Harvey
Manuscript details:
Ms
# o2767
Received
19 April 2011
Final
received 10 September 2011
Finally
accepted 18 October 2011
Citation: Hu, J. & F. Zhang (2011). Description of
three new species of the genus Allochthonius Chamberlin, 1929
(Pseudoscorpiones: Pseudotyrannochthoniidae) from China. Journal of Threatened Taxa 3(11): 2167-2176.
Copyright: © Junfang Hu & Feng Zhang 2011. Creative
Commons Attribution 3.0 Unported License. JoTT allows unrestricted use
of this article in any medium for non-profit purposes, reproduction and
distribution by providing adequate credit to the authors and the source of
publication.
Author
details: Junfang
Hu is a master student in Hebei
University, Hebei Province, China. Her major is zoology, mainly to classify the pseudoscorpions on fauna of
China, and her supervisor is Prof. Feng Zhang.
Prof. Feng Zhangis currently teaching students, undertaking and supervising research activities
in Hebei University. His recent
research focus is on systematics of Arachnida. He has published more than 90
research papers in leading national and international journals. He has produced
12 master students in field of Arachnida.
Author Contribution: JH undertook illustration, identification and wrote the
manuscript. FZ supervised Junfang
Hu during the study and provided necessary suggestions during manuscript
preparation.
Acknowledgements: This text benefited greatly from helpful comments by Dr. Volker
Mahnert (Muséum d’Histoire Naturelle, Geneva), and many thanks are expressed to
Dr. Mark Harvey (Western Australian Museum, Perth, Australia) for supplying
data for inclusion in the map. Thanks are also given to Dr. Hiroshi Sakayori
(Mitsukaido Daini Senior High School) and Dr. Hidebumi Sato (Tsurumi Girls’
Senior High School) for kindly donating valuable literature. This work was
supported by the National Natural Science Foundation of China (No.
31071885, 30970325, 31093430).
Abstract: The pseudoscorpion genus Allochthonius Chamberlin, 1929, belonging to the family
Pseudotyrannochthoniidae Beier, 1932, is reported from China, and the
subgeneric characters of Allochthonius (Allochthonius) are reviewed in detail. Three new species are diagnosed,
described and illustrated under the names Allochthonius (A.)fuscus sp. nov., A. (A.) wui sp. nov. and A. (A.) trigonus sp. nov. A
distribution map and a key to the species of subgenus Allochthonius (A.)
are provided. In addition, Centrochthonius sichuanensisSchawaller, 1995
is transferred to Allochthonius, forming the new combination A. (A.) sichuanensis (Schawaller).
Keywords: Allochthonius, Centrochthonius, China, new species, pseudoscorpions, taxonomy, transfer.
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INTRODUCTION
The pseudoscorpion genus Allochthonius was erected by Chamberlin (1929) for the Japanese
type species Chthonius opticus Ellingsen, 1907. This genus was later divided into three subgenera, Allochthonius, Urochthonius Morikawa, 1954 and Spelaeochthonius Morikawa, 1954, by
Morikawa (1960) based on the presence or absence of eyes, carapacal chaetotaxy
and characteristics of the cheliceral fixed finger teeth. Muchmore (1967) used the morphology of
the coxal spines and carapacal chaetotaxy to transfer Allochthonius (Spelaeochthonius) to PseudotyrannochthoniusBeier,
1930. Harvey (1991, 2009) agreed
with this view which is reflected in the catalogue of the Pseudoscorpiones, and
which we follow in this paper. Presently, the genus Allochthonius is composed of two subgenera Allochthonius and Urochthonius, which are widely
distributed in Japan and South Korea. The subgenus Allochthonius (Allochthonius) is composed of seven species, and Allochthonius (Urochthonius) includes eight species
(Harvey 2009).
While examining pseudoscorpion specimens collected by Dr. Min Wu
and Prof. Fusheng Huang from southern and western China, we found some Allochthonius specimens belonging to the
subgenus Allochthonius (Allochthonius). Three new species
are recognized, which are described and illustrated in this paper. In addition, we assess the taxonomic
position of Centrochthonius sichuanensis Schawaller, 1995 whichhas many characteristics of Allochthonius.
MATERIALS AND METHODS
The patterns of description and the terminology follow Chamberlin
(1931) and Harvey (1992). This
setal formula of the palpal femur mainly follows Vachon (1941), but the
difference from his method is that we divided the “ventral” to “posteroventral”
and “anteroventral” (rows in sequence anterior-anterodorsal-dorsal-posterodorsal-posterior-postero-ventral-anteroventral). The term “rallum” (for flagellum) is
adopted following Judson (2007). All specimens are preserved in 75% alcohol and were examined and
illustrated under a Leica M165c stereo-microscope with
a drawing tube, which was also used for the measurements. Detailed examination was carried out
with a Nikon YS100 general optical microscope. Temporary slide mounts were made in glycerol. All measurements are given in mm. The specimens referred here are
deposited in the Museum of Hebei University (MHBU), Baoding City, China.
The following abbreviations are used in the text for the
trichobothria: b - basal; sb - sub-basal; st - sub-terminal; t - terminal; ib -
interior basal; isb - interior sub-basal; ist - interior sub-terminal; it -
interior terminal; eb - exterior basal; esb - exterior sub-basal; est -
exterior sub-terminal; et - exterior terminal; dx -trichobothria dx. 6–4,
18 refer to carapacal chaetotaxy: carapace with 18 setae, anterior margin with
six setae and posterior margin with four setae.
RESULTS
Allochthonius (Allochthonius) Chamberlin, 1929
Morikawa, 1960: 98; Harvey, 1991: 132.
Description: Carapacal chaetotaxy: 6–4, 18; 8–4, 22; 8–4,
24; 10–4, 24; 10–4, 26; 10–4, 28; or 10–4, 30. Four eyes, anterior eyes with developed
tapeta and situated on eye-tubercles, posterior eyes with less developed tapeta
than anterior ones and without eye-tubercles. Epistome and spinneret are generally absent but sometimes
present. Cheliceral palm with five
or six setae, fixed finger generally provided with one large basal and one
subapical teeth, and between them two or three small teeth (or with only one
basal large tooth and a few small teeth before it, or with several small teeth
on the median swelling without any large tooth). Rallum composed of a biseriate row of about eight to eleven
pinnate setae. Chelal
finger with well-spaced and prominent marginal teeth, and movable finger with a
tubercle between two teeth (Figs. 6, 13, 21, 26). Coxal spines present
on coxae I only, consisting of a tubercle expanded terminally into a spray of
about four to ten clavate, fan-shaped or gladiate spines. With a well developed bisetose
intercoxal tubercle. Palps, chelicerae and legs relatively short and robust.
Distribution: China, Japan, South Korea.
Remarks: Allochthonius (Allochthonius) is similar to the subgenus Allochthonius (Urochthonius) in that: the chelal
fingers have well-spaced and prominent marginal teeth; coxal spines are only
present on coxae I, consisting of a tubercle expanded terminally into a spray
of about eight processes, which extend anteriorly and more or less shield the
apical process of coxae I (Morikawa 1960). The two subgenera A. (Allochthonius) and A. (Urochthonius) can be distinguished by the presence of four eyes, or blind or
rarely with two eyes, respectively, which is consistent with Morikawa’s (1960)
viewpoint, but the differences from his system is by the dental morphology and
tooth number on the cheliceral fixed finger and the number of setae on the
cheliceral palm. For instance, Allochthonius (A.) montanus and Allochthonius (A.) shintoisticus both have four eyes, but
the cheliceral fixed finger has one large and a few small teeth and five setae
on the cheliceral palm in the former, and the cheliceral palm with five setae
in the latter.
In addition, while examining the specimens collected from China,
we found a significant characteristic in all species: the palpal movable finger
has a tubercle between two teeth. This tubercle is present in Allochthonius (A.) tamurai (cf. Sakayori, 2003: 25,
fig. 5a; 27, fig. 21a), but Sakayori (2003) did not refer to this tubercle in
the description. We infer that this characteristic might only exist in the
subgenus Allochthonius (Allochthonius), so further study to all species of Allochthonius, with especial attention
to the movable chelal fingers, is required.
Composition: The subgenus Allochthonius (Allochthonius) comprises seven species (including two subspecies), and
distributed in East Asia (Fig. 27); five species distributed in Japan (Allochthonius (A.) borealis Sato, 1984, Allochthonius (A.) montanus Sakayori, 2000, Allochthonius (A.) shintoisticus Chamberlin, 1929, Allochthonius (A.) tamurai Sakayori, 1999, Allochthonius (A.) opticus (Ellingsen, 1907) (with the
subspecies Allochthonius (A.) opticus opticus (Ellingsen, 1907), Allochthonius (A.) opticus troglophilus Morikawa, 1956); two species, Allochthonius (A.) buanensis W.K. Lee, 1982 and Allochthonius (A.) coreanus Morikawa, 1970, are in
South Korea. The three new species
described here also belong to this subgenus (Allochthonius (A.) trigonus sp. nov., Allochthonius(A.) fuscus sp. nov. and Allochthonius (A.) wui sp. nov.). In addition, we
also include Allochthonius (A.) sichuanensis (Schawaller, 1995) from China in this subgenus.
Allochthonius
(Allochthonius) sichuanensis (Schawaller, 1995),comb. nov.
Centrochthonius
sichuanensis Schawaller, 1995: 1046-1048, Figs. 1–5.
Remarks: Centrochthonius sichuanensis Schawaller, 1995 was described from material
collected in Wolong Nature Reserve, Sichuan Province, China. The description indicates that the
epistome is absent, the coxal spines are on a tubercle, and the carapace has a
high number of setae. All of these
characters conform to the diagnosis of Allochthonius, not Centrochthonius (M.S. Harvey, pers.
comm.). The presence of eyes
suggests that it can be treated in the subgenus Allochthonius. We therefore transfer this species to Allochthonius (Allochthonius), and form a new
combination, Allochthonius (A.) sichuanensis (Schawaller, 1995).
Allochthonius (Allochthonius) fuscus sp. nov.
(Figs. 1–8)
Type material: Holotype male,
24.iii.1975, 26004’N & 119021’E, Fuzhou City, Fujian
Province, China, (Ps.-MHBU-FJ750324), Fusheng Huang leg.;Paratypes: 03.iii.1983, one male (Ps.-MHBU-FJ750325), two females
(Ps.-MHBU-FJ750326–750327), 30013’N & 120006’E,
alt. 134m; Hangzhou City, Zhejiang Province, China, Fusheng Huang leg.
Diagnosis: Chelal palm obviously dark
in color; carapacal chaetotaxy (Fig. 1) 8–4–4–2–4 (22);
coxal spines present on coxae I and consisting of 10 tridentate blades, each
blade with a central spine terminally distinctly expanded as fan-shaped, all
situated on a common tubercle (Fig. 4). This new species is closely related to Allochthonius (A.) tamurai from Japan, but can be
distinguished from the latter by the number of coxal spines (six blades spines
in latter) and by the shape of the coxal spines (gladiate in latter).
Etymology: The specific name is
derived from the Latin word “fuscus” means dark colored, referring to the color
of the chelal palm.
Description: Relatively large species. Body (Image 1) yellow, chelal palm strong yellow and other segments
of chela pale yellow. Carapace
subquadrate and slightly shorter than broad (0.9 times), carapace indistinctly
constricted posteriorly; anterior eyes with well developed tapeta and situated
on tubercles, posterior eyes less developed tapeta than anterior ones and
without eye-tubercles; epistome absent, space between median setae straight or
slightly recurved; chaetotaxy 8–4–4–2–4 (22).
Tergal chaetotaxy 4: 4–7: 6–9: 7–9:
7–10: 8–11: 10–11: 10–12: 8–10: 7–10: 2–4
(two tactile setae): 0. Male anterior genital operculum with eight setae,
genital opening pit-like in the basal half, 11 marginal setae on each side;
sternal chaetotaxy: 12–13 + 2×2–3 suprastigmatic microsetae:
12–13 + 2×2–3: 15: 15: 12: 13: 12: 9: 0: 2. Coxae typical,
setae P 5, I 4, II 5, III 7, IV 7; intercoxal tubercle present with two setae.
Coxae I each with ten spines, arranged on a common tubercle.
Cheliceral palm with six setae, of
which a minute one is located laterally; palm with moderate hispid granulation
interiorly and laterally. Fixed finger of
holotype (Fig. 2) with six teeth, of which nearly equal length (paratype (Fig.
3) with one large basal and one subapical teeth, between them with two small
teeth); movable finger with 14 relatively small teeth of equal length; spinneret
absent. Serrula exterior with 17 lamellae, serrula interior
with 13 lamellae. Rallum composed of 11 blades with fine barbules, of
which the posterior blade is shorter than others.
Palp smooth, femur 2.0 times longer than
carapace, setal formula 7–9–4–3–5–5–2;
chelal palm distinctly expanded towards internal side, chelal finger straight in
dorsal view; fixed finger with 17 teeth, first three and last four teeth smaller
than others; movable finger with 18 teeth, all of which nearly equal length and
smaller than fixed finger teeth, and with a tubercle between the ninth and
tenth teeth from distal end (Fig. 6).
Legs (Figs. 7, 8) typical. Femur of leg I 2.2
times longer than patella, tarsus 2.0 times longer than tibia, setae of leg I
(trochanter to tibia) 6: 15: 12: 10, setae of leg IV (trochanter to metatarsus)
3: 2: 9: 15: 12; patellae of legs III and IV each with four setae in dorsal
row; femur III and IV without dorsal setae; trochanter II with four setae,
trochanter III with two setae. Leg
IV with two tactile setae present on metatarsus (TS= 0.27) and tarsus (TS=
0.23).
Measurements (ratios in parentheses; for
the palp, the larger measurements and lower ratios only referring to male). Body length 2.25. Carapace 0.49×0.54
(0.9). Chelicera 0.52×0.26 (2.0), movable finger
length 0.30. Palp femur 1.05–1.22×0.20–0.22
(5.3–5.5), patella 0.50–0.55×0.21–0.23 (2.3–2.4),
chela 1.47–1.80×0.30–0.36 (4.9–5.0), palm length 0.52–0.69
(1.7–1.9), movable finger length 0.94–1.02 (1.5–1.8 ×
palm). Leg I
femur 0.47×0.08 (5.9), patella 0.29×0.07 (4.1), tibia 0.28×0.06
(4.7), tarsus 0.46×0.05 (9.2); leg IV femur+patella 0.75×0.23
(3.3), tibia 0.57×0.10 (5.7), metatarsus 0.26×0.08 (3.3), tarsus
0.52×0.05 (10.4).
Distribution: China (Fujian, Zhejiang).
Allochthonius (Allochthonius) wui sp. nov.
(Figs. 9–16)
Type material: Holotype male, 07.vi.1997,
36050’N &101059’E, alt. 2699m, Huzhu National Forest
Park, Qinghai Province, China, (Ps.-MHBU-QH970607), Min Wu leg.; Paratypes: 08.vi.1997, one male (Ps.-MHBU-QH970608), 12
deutonymphs (Ps.-MHBU-QH970609–970620), 36050’N & 101059’
E, alt. 2699m, Huzhu National Forest Park, Qinghai Province, China, Min Wu leg.
Diagnosis: Large species and body
with dark color (Image 2); coxal spines present on coxae I
and consisting of ten tridentate blades, each spine with the central one
terminal distinctly fan-shaped (Fig. 11). This species resembles Allochthonius (A.) shintoisticus Chamberlin, 1929 from Japan, but differs from it by the
chaetotaxy of the carapace (22 setae in A. wui and 24 in A. shintoisticus), by the cheliceral palm
(six setae in A. wui and five in A. shintoisticus), and by the shape of coxal spines (fan-shaped in A. wui and clavate in A. shintoisticus).
Etymology: The specific name is named
after Prof. Min Wu, who collected the specimens.
Description: Body deep brown, and chelicerae
dark yellow. Carapace subquadrate
(0.9 times), basally constricted, chaetotaxy 8–4–4–2–4,
22; epistome absent; anterior eyes with well developed tapeta and situated on
eye-tubercles, posterior eyes with less developed tapeta than anterior ones and
without eye-tubercles.
Tergal chaetotaxy 4: 5–6: 6: 7: 9: 10: 10–11:
10–11: 8: 6: TT: 0. Male anterior genital operculum with eight setae,
genital opening pit-like in the basal half, 12 marginal setae on each side;
sternal chaetotaxy: 12 + 2×3 suprastigmatic microsetae: 12 + 2×3:
13: 12: 13: 13: 12: 8: 0: 2. Coxae typical, setae P 5, I 4, II 5, III 6, IV 6;
intercoxal tubercle present with two setae. Holotype male coxae I each with 10
tridentate spines (paratype male with nine spines), nine blades arranged on a
common tubercle, one blade aside.
Cheliceral palm with six setae, of which a
minute one is located laterally; palm nearly smooth, but near the base of fixed
finger and interior side with distinctly acute granules (Fig. 10); right fixed finger
with one large basal and subapical tooth and between them with three small
teeth, but the left finger with two large basal teeth and one large subapical
tooth, between them two small teeth; movable finger with 13 small teeth of equal
length; spinneret absent, serrula exterior with 19 lamellae, serrula interior
with 16 lamellae. Rallum composed of 11 dentate blades, of which the posterior
one is smaller than others.
Palpal femur 2.0 times longer than carapace,
setal formula 11–10–5–5–7–8–3; chelal palm
distinctly expanded towards internal side, chelal finger straight in dorsal
view; fixed finger with 18 acute teeth; movable finger with 21 teeth, and with a
tubercle between the seventh and eighth teeth from terminal (Fig. 14).
Legs (Figs. 15, 16) typical. Femur 1.7 times
longer than patella, patella 3.7 times longer than deep, tarsus 1.9 times
longer than tibia, setae of leg I (trochanter to tibia) 5: 11: 10: 11, setae of
leg IV (trochanter to metatarsus) 2: 2: 9: 13: 14; patellae of legs III and IV
each with four setae in dorsal row; femur III and IV without dorsal setae;
trochanter II with five setae, trochanter III with three setae. Leg IV
metatarsus and tarsus each with one basal tactile seta, tactile setae present
on metatarsus (TS= 0.22) and tarsus (TS= 0.24).
Measurements (ratios in
parentheses; for the palp, the larger measurements and lower ratios only
referring to male). Body
length 2.50. Carapace 0.52×0.58 (0.9). Chelicera 0.55×0.25 (2.2), movable finger length 0.30. Palp femur 1.36–1.40×0.19–0.23
(6.1–7.2), patella 0.52–0.58×0.21–0.23 (2.4–2.5),
chela 1.90–2.03×0.39–0.40 (4.9–5.1), palm length 0.66–0.73
(1.7–1.8), movable finger length 1.13–1.25 (1.7 × palm).Leg I femur 0.57×0.08 (7.1), patella 0.37×0.08 (4.6), tibia 0.30×0.07
(4.3), tarsus 0.58×0.06 (9.7); leg IV femur+patella 0.92×0.22
(4.2), tibia 0.65×0.12 (5.4), basitarsus 0.32×0.08 (4.0),
telotarsus 0.67×0.06 (11.2).
Distribution: China (Qinghai).
Allochthonius (Allochthonius) trigonus sp. nov.
(Figs. 17-23)
Type material: Holotype male, 03.v.1978,
27040’N & 118002’E, alt. 200m, Chong’an City, Fujian
Province, China, (Ps.-MHBU-FJ780503), Fusheng Huang leg.
Diagnosis: Anterior margin of carapace
with 27 triangular protuberances of which eight are situated near the anterior
eyes (Fig. 17); coxal spines present on coxae I and consisting of seven
tridentate blades, each blade with a central branch terminally distinctly expanded
as fan-shaped, six spines on a common tubercle and a single spine slightly
separate (Fig. 19). This present
species resembles Allochthonius (A.) borealis, but is distinguishable from the latter by the number of blades
in the rallum (11 blades, in A. trigonus, and eight blades in A. borealis) and the epistome (present
in latter).
Etymology: The specific name is
derived from the Greek word “trigonon”, means triangle, referring to the shape
of triangular protuberances on the carapace anterior margin.
Description: Moderately large species. Body bright yellow. Carapace subquadrate and slightly
constricted posteriorly, shorter than broad (0.8 times), carapace anterior
margin with 27 triangular protuberances of which eight near the anterior eyes;
anterior eyes with developed tapeta and situated on eye-tubercles, posterior
eyes with less developed tapeta than anterior ones and without eye-tubercles;
epistome absent, space between median setae straight or slightly recurved;
chaetotaxy 10–4–6–2–4 (26).
Tergal chaetotaxy 4: 4: 7: 6: 8: 10: 9: 12: 11:
8: T2T: 0. Male anterior genital operculum with eight setae, genital opening
pit-like in the basal half, 13 marginal setae on each side; sternal chaetotaxy:
14 + 2×3 suprastigmatic microsetae: 14 + 2×3: 14 + 2×3: 13:
13: 12: 12: 11: 11: 0: 2. Coxae
typical, setae P 5, I 4, II 5, III 6, IV 6; intercoxal tubercle present with
two setae. Coxae I each with seven spines, of which six arranged on a common
tubercle and a single spine aside.
Chelicera palm with six setae, of which a minute
one located laterally; palm nearly smooth, but near the base of fixed finger
with distinct granules (Fig. 18), fixed finger with one large basal and
one subapical teeth, between them with two small teeth inserted; movable finger
with 18 small teeth of equal length; spinneret absent. Serrula exterior with
17 lamellae, serrula interior with 14 lamellae. Rallum composed of 11
dentate blades, of which the posterior one smaller than others.
Palp smooth, apart from a
patch of minute denticles on anterior surface of trochanter (Fig. 20). Femur 1.7 times longer
than carapace, setal formula 8–11–4–4–5–2–1;
chelal palm distinctly expanded towards internal side; chelal finger straight in
dorsal view; chelal fixed finger with 18 teeth, of which the basal two smaller
than others; movable finger with 16 teeth, with a tubercle between the sixth and
seventh teeth from terminal (Fig. 21).
Legs (Figs. 22, 23) typical. Femur of leg I 1.6
times longer than patella, tarsus 2.0 times longer than tibia. Setae of leg I
(trochanter to tibia) 3: 12: 11: 8, setae of leg IV (trochanter to metatarsus)
2: 2: 9: 12: 12; patellae of legs III and IV each with four setae in dorsal
row; femur III and IV without dorsal setae; trochanter II with four setae,
trochanter III with four setae. Leg IV with two tactile setae present on metatarsus (TS= 0.33) and
tarsus (TS= 0.23).
Measurements. (ratios in
parentheses). Body length 2.0. Carapace
0.43×0.51 (0.8). Chelicera 0.48×0.22
(2.2), movable finger length 0.26. Palp femur 0.80×0.16
(5.0), patella 0.35×0.15 (2.3), length of chela 1.22×0.23 (5.3),
palm length 0.39 (1.7), movable finger length 0.82 (2.1 × palm).Leg I femur 0.42×0.08 (5.3), patella 0.24×0.08 (3.0), tibia 0.23×0.08
(2.9), tarsus 0.43×0.05 (8.6). Leg IV femur+patella 0.68×0.22
(3.1), tibia 0.49×0.10 (4.9), metatarsus 0.23×0.08 (2.9), tarsus 0.50×0.05 (10.0).
Distribution: China (Fujian).
Remarks: Allochthonius (A.) trigonus sp. nov. is distinguished from the two
previously described species Allochthonius (A.) fuscus sp. nov. and Allochthonius (A.) wui sp. nov. by the triangular protuberances
present on anterior margin of carapace (absent in the other species) and the
slender chela (movable finger 2.1 × palm vs 1.5–1.8 × palm). Allochthonius (A.) fuscus sp. nov. shares with Allochthonius (A.) wui sp. nov. the same carapacal chaetotaxy
(8–4–4–2–4 (22)), but differs from it by a slightly
stouter palpal femur (5.3–5.5 times vs 6.1–7.2 times longer than
broad) and chela (1.47–1.80 mm vs 1.90–2.03 mm).
Allochthonius (A.) sp.
(Figs. 24–26)
Material examined: Female, 04.vii.1990, 32037’N& 103036’E, Ngawa, Songpan County, Sichuan Province, China,
(Museum ID # Ps.-MHBU-SC900704), Fusheng Huang leg.
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