Journal of Threatened Taxa | www.threatenedtaxa.org | 26
November 2019 | 11(14): 14808–14815
Barilius torsai (Teleostei: Cypriniformes: Cyprinidae), a new freshwater fish from the Brahmaputra
drainage, India
Kavita Kumari
1, Manas Hoshalli Munivenkatappa 2, Archana Sinha 3, Simanku Borah 4 & Basanta
Kumar Das 5
1,2,3,5 ICAR-Central Inland Fisheries Research Institute, Monirampur (Post), Barrackpore, Kolkata, West Bengal
700120, India.
2 ICAR-Central
Marine Fisheries Research Institute Regional Centre, Andhra University (Post),
Opposite SBI Kohinoor Branch, Vishakapatnam, Andhra
Pradesh 530003, India.
4 ICAR-CIFRI
Centre, HOUSEFEED Complex, Dispur (Last gate), Guwahati, Assam 781006, India.
1 kavitacof@gmail.com,
2 manas2u@gmail.com, 3 sinhaarchana@yahoo.com,
4 simankuborah@gmail.com
(corresponding author), 5 basantakumard@gmail.com
Abstract: Barilius torsai is described from the Torsa,
a tributary of Brahmaputra River system in West Bengal, India. The new species
is distinguished from all its congeners by the presence of a complete lateral
line with 52–53 scales, 29 pre-dorsal scales, pectoral fin notched, two
well-developed pairs of barbels (rostral and
maxillary), length of rostral barbel slightly larger
than maxillary, which reaches the orbit.
Tubercles on snout and lower jaw absent, 9–11 blue vertical bars along
the body, dorsal fin hyaline with dark pigment concentrated along lower
two-third of the dorsal-fin rays.
Keywords: Chedrini, Danioninae, taxonomy, Torsa, West Bengal.
doi: https://doi.org/10.11609/jott.4746.11.14.14808-14815 | ZooBank: urn:lsid:zoobank.org:pub:591E7FA3-82AC-48DE-B249-F09BBC477BE2
Editor:
Rajeev Raghavan, Kerala
University of Fisheries and Ocean studies (KUFOS), Cochin, India. Date of
publication: 26 November 2019 (online & print)
Manuscript
details: #4746 | Received 05
December 2018 | Final received 09 September 2019 | Finally accepted 12 November
2019
Citation: Kumari, K., M.H. Munivenkatappa,
A. Sinha, S. Borah & B.K. Das (2019).
Barilius torsai (Teleostei: Cypriniformes: Cyprinidae), a new
freshwater fish from the Brahmaputra drainage, India. Journal
of Threatened Taxa
11(14): 14808–14815. https://doi.org/10.11609/jott.4746.11.14.14808-14815
Copyright: © Kumari et al. 2019. Creative Commons Attribution 4.0 International
License. JoTT
allows unrestricted use, reproduction, and distribution of this article in any
medium by adequate credit to the author(s) and the source of publication.
Funding: Indian Council of Agricultural Research under the Project-REF/ER/12/01/02.
Competing interests: The authors declare no competing interests.
Author
details: Kavita Kumari is scientist,
Fish Genetics and Breeding at ICAR-CIFRI, Barrackpore and working on taxonomy,
systematics and stock characterisation of fish along
with assessment of ecosystem health using molecular tools. Manas Hoshalli Munivenkatappa is
scientist at ICAR-CMFRI and working on marine fisheries management, stock
assessment, fish biology and Fisheries policy. Archana Sinha is Principal scientist at ICAR-CIFRI and
working on various aspects of small indigenous fish culture and
conservation. Simanku Borah is scientist at ICAR-CIFRI Regional Centre, Guwahati
and working in the field of Fishery Biology. He is also involved in research
related to ecology and habitat characterization of rivers and wetlands. Basanta Kumar Das is Director, ICAR-CIFRI,
Barrackpore and working on the aspects of Inland Fisheries Management.
Author
contribution: KK—collected and analyzed specimens, examined the
museum specimens, wrote final version of manuscript. MHM— analyzed specimens
and helped in the manuscript preparation.
AS—supervised study, helped in the revision of the manuscript,
SB—examined the museum specimens, helped in revision of the manuscript.
BKD—supervised study, helped in the revision of the manuscript.
Acknowledgements: The authors are very
grateful to Prof. A. P. Sharma, Former Director,
ICAR-CIFRI and Dr. V. R. Suresh, Head of Division,
Riverine Ecology and Fisheries for providing facilities for the work. We also
thank Director, ZSI, Kolkata, Dr. L. Kosygin Singh,
In-charge Freshwater Fish Section, Indian Museum, ZSI, Kolkata, Dr. Bikramjit Sinha,
Officer-in-Charge, ZSI, Arunachal Pradesh, Dr. W.
Vishwanath Singh, Professor, Department of Life Science, Manipur University and
Dr Rameshori Yumnam,
Assistant Professor, Department of Life Science, Manipur University for
necessary permission and help to examine comparative material. The authors also acknowledge Dr. M.K. Mukhopadhyay retired scientist, ICAR-CIFRI for his
guidance for the study.
Introduction
Fishes of the genus Barilius
Hamilton (Cyprinidae:Danioninae: Chedrini)
are one of the dominant small-sized fishes occurring in hill streams and upland
rivers throughout India, Bangladesh, Myanmar, Nepal, Sri Lanka, China, Iraq,
and Thailand (Selim & Vishwanath 2002; Jayram 2010; Tejavej
2012; Qin et al. 2019). Barilius was erected as a subgenus of Cyprinus with Cyprinus
barila Hamilton, as the type (Qin et al.
2019). Howes (1980) identified two
groups within Barilius (sensu
lato), corresponding to the genera Barilius and Opsarius
(Rainboth 1991).
These bariliine fishes are characterised
by their relatively elongate, compressed bodies with round belly, vertical bars
on the flanks, 9–17 anal-fin rays and sub-laterally placed lateral line
(Hamilton 1822; Howes 1980; Talwar & Jhingran
1991; Tejavej 2012).
Of the 36 valid species of bariliine fishes,
24 have been recorded in India (Fricke et al. 2019; Qin et al. 2019). Currently, most bariliine
fishes, including all southeastern Asian species are assigned to the genus Opsarius (Howes 1983; Rainboth
1991; Qin et al. 2019) and the genus Barilius
is restricted to five species characterized by an extremely shallow body: B.
barila (Hamilton, 1822), B. evezardi Day, 1872, B. modestus
Day, 1872 and B. vagra (Hamilton, 1822) from
India, and B. mesopotamicus Berg, 1932 from
the Tigris-Euphrates basin, based on the diagnosis of Barilius
sensu by Howes (1980), as well as morphological
(Howes 1980; 1983) and molecular phylogenetic evidence (Tang et al. 2010; Liao
et al. 2011; Qin et al. 2019).
During an ichthyological survey in the Torsa, a tributary of the Brahmaputra River during July and
November 2015, a total of 83 individuals of a Barilius
species were caught by cast net of mesh size 15–20 mm at Jaldapara, Alipurduar District,
West Bengal, India. Examination of the
collected specimens in July 2015 and detailed re-examination in November 2015
following standard literature (Jayaram 2010; Arunkumar
& Singh 2000; Nath et al. 2010; Dishma &
Vishwanath 2012; Knight et al. 2015) revealed that 11 of the 83 specimens could
not be assigned to any of the known species.
The new species is described herein as Barilius
torsai.
Materials and Methods
Measurements were made with digital caliper with an accuracy of 0.1mm. Counts and measurements were made on the left
side of the specimens wherever possible and based on standard methods following
Dishma & Vishwanath (2012). Colour pattern was recorded from fresh and
preserved (10% formalin) specimens. Voucher specimens are deposited in the
Museum of the Zoological Survey of India (ZSI), Kolkata and at the ICAR-Central
Inland Fisheries Research Institute (CIFRI), Barrackpore.
Barilius torsai sp. nov.
(Image 1)
urn:lsid:zoobank.org:act:79FEC835-A4FA-4D27-8EC1-2EDD65616A31
Type material
Holotype: ZSI FF5542, 12.xi.2015, 26.7290N
& 89.3250E, 71.41mm SL, Torsa River, Jaldapara, Alipurduar District,
West Bengal, India, coll. A. Roy Chaudhary.
Paratype: ZSI FF5543, 26.vii.2015, 26.7290N
& 89.3250E, 74.56mm SL, data same as for holotype, coll. A. Mitra; CIFRI F10003-10010, 8 ex., 26.vii.2015, 71.46–74.23
mm SL, data same as for holotype, coll. A. Mitra;
CIFRI F10011, 12.xi.2015, 71.46mm SL, data same as for holotype, coll. A. Roy
Chaudhary.
Diagnosis
Barilius torsai is distinguished from all other species of Barilius by a combination of the following
characters: lateral line complete with 52–53 scales, 29 pre-dorsal scales,
pectoral fin notched, two well-developed pairs of barbels
(rostral and maxillary), body with 9–11 blue vertical bars, dorsal fin hyaline
with dark pigment concentrated along lower 2/3rd of dorsal-fin rays.
Description
See Table 1 and Supplementary 1 for morphometric
characters and image 1 for general appearance.
Body shallow, its depth about one-fourth standard length (SL), laterally
compressed with ventral profile more convex than the dorsal profile. Caudal peduncle long, narrower near the
caudal base. Head small and compressed,
length about one-fourth SL, snout slightly blunt. Mouth oblique, angle of gape not reaching
vertical from the anterior margin of the orbit.
No symphysial process in the lower jaw.
Eyes large, situated in the anterior half of the head, diameter about
one-fourth head length (HL). Nostrils
closer to anterior margin of eye than the snout tip. Two pairs of well-developed barbels (rostral and maxillary), length of rostral barbel slightly larger than maxillary, which reaches the
orbit. Tubercles on snout and lower jaw
absent.
Dorsal fin with two simple and seven branched rays,
its origin posterior to the pelvic-fin origin, and closer to the caudal-fin
base than tip of the snout. Pectoral fin
with two simple and 11 branched rays, not reaching the pelvic-fin base, notched
after third (between third and sixth ray).
Pelvic fin with two simple and seven branched rays, its origin closer to
the anal-fin than pectoral-fin origin, posterior tip not reaching anus/anal fin
origin, muscular pad at base present.
Anal fin with two simple and eight branched rays, its origin just below
base of the last dorsal fin ray. Caudal
fin deeply forked, lobes equal, with 17 principal rays. Scales cycloid and small. Lateral line
complete, slightly curved, running along lower half of the body and passing
almost through middle of the caudal base, with 52–53 scales in the lateral-line
row up to the end of the caudal base.
Scales in transverse line on body 11/1/5 between dorsal fin
origin and pelvic fin base. 29
pre-dorsal scales.
Colour
In live and fresh specimens, dorsum appears
greyish, sides and belly silvery. Body
with 9–11 blue vertical bars, three to five anterior bars almost reaching
lateral line, number of bars on either side of the body unequal. Pectoral, pelvic and anal fin hyaline. Caudal fin hyaline, with black margin on fork
edges. Dorsal fin hyaline with dark
pigment on rays, concentrated along lower two-third of dorsal-fin rays. In preserved specimens, silvery colouration
disappears and all dark pigment in fins and body bar turns black. Dorsum appears black and ventral areas creamy.
Etymology
The specific name refers to the Torsa, type locality of the species, and a tributary of the
Brahmaputra River System. An adjective.
Distribution
Presently known only from the Torsa
River in West Bengal, India (Figure 1, Image 2).
Discussion
Fifteen species of bariliine
fish are found in the Brahmaputra basin, and the larger northeastern
Indian region. They are Barilius vagra (Hamilton,
1822); B. barila (Hamilton, 1822); Opsarius arunachalensis
(Nath et al., 2010); O. bendelisis
(Hamilton, 1807); O. howesi
(Barman, 1986); O. lairokensis (Arunkumar & T. Singh, 2000); O. profundus (Mayanglambam
& Vishwanath, 2012); O. radiolatus
(Gunther, 1868); O. Shacra (Hamilton,
1822); O. barna (Hamilton, 1822); O. chatricensis (Selim & Vishwanath, 2002); O. dimorphicus (Tilak & Husain, 1990); O. dogarsinghi (Hora, 1921); O. ngawa
(Vishwanath & Manojkumar, 2002), and
O. tileo (Hamilton, 1822). Barilius torsai sp. nov. is a member
of Barilius. It can be distinguished from members of Opsarius sensu
Howes (1980; 1983) in having a shallow body (vs. deep), two pairs of barbels (vs. a single pair or absent), elongated pelvic
axial scales (vs. lobate or fleshy), and absence of parallel rows of tubercles
on the dentary (vs. presence). Barilius torsai can
be distinguished from all other congeners by the presence of notched pectoral
fin, and from its northeastern Indian congeners by
the presence of a complete lateral line with 52–53 scales, greater number of
pre-dorsal scales and hyaline dorsal fin with dark pigment concentrated along
lower two-third of the dorsal-fin rays and lesser body depth at dorsal-fin
origin except B. barila, O. shacra, O. bendelisis,
O. chatricensis (Table 2). Barilius torsai further differs from species of B. vagra and B. barila in
having greater number of predorsal scales (29 vs.
21–22 in B. vagra and 22 in B. barila), lateral line scales (52–53 vs. 40–45 in B. vagra and B. barila)
and lesser number of branched pectoral fin rays (11 vs. 14–15 in B. vagra and 12 in B. barila).
In comparison to the species currently included within
Opsarius sensu
Howes (1980; 1983) Barilius torsai is similar to O. shacra
and O. arunachalensis
in its dorsal and anal fin ray counts, but differs from O. shacra in having
lesser number of lateral line scales (52–53 vs. 59–70), greater number of
pre-dorsal scales (29 vs. 22–25), branched pectoral fin rays (11 vs. 14) and
the dorsal fin with dark pigment along lower the two-third margin (vs. upper
third). It differs from O. arunachalensis in having a greater body depth at
dorsal-fin origin (22.17–23.89% SL vs. 20.21–20.83% SL), greater number of
lateral line scales (52–53 vs. 40–45) and presence (vs. absence) of barbels and vertical bars on body.
Barilius torsai differs from O. profundus
and O. lairokensis in having lesser body
depth (22.17–23.89% SL vs. 32.1–37.4% SL in O. profundus
and 25.54% SL in O. lairokensis),
greater number of lateral line scales (52–53 vs. 30–35 in O. profundus and 41–44 in O. lairokensis)
and greater number of predorsal scales (29 vs. 17–18
in O. profundus and 21 in O. lairokensis). Barilius
torsai further differs from O. bendelisis, O. howesi and O.
radiolatus in having greater number of predorsal scales (29 vs. 18–20 in O. bendelisis,
17–18 in O. howesi and 24-25 in O. radiolatus), lateral line scales (52–53 vs. 39–46 in O.
bendelisis, 42–43 in O. howesi
and 56–62 in O. radiolatus) and
lesser number of branched pectoral fin rays (11 vs. 14 in O. bendelisis, 13 in O. howesi
and 16 in O. radiolatus).
Barilius torsai differs from O. barna,
O. chatricensis, O. dimorphicus,
O. dogarsinghi, O. ngawa,
and O. tileo in having greater number of predorsal scales (29 vs. 15–16 in O. barna,
15 in O. chatricensis, 25–27 in O.
dimorphicus, 20 in O. dogarsinghi,
21–22 in O. ngawa, and 28 in O. tileo), lateral line scales (52–53 vs. 36–42 in O. barna, 38 in O. chatricensis,
60–66 in O. dimorphicus, 37–40 in O. dogarsinghi, 40–43 in O. ngawa,
and 59+4 in O. tileo) and lesser number of
branched anal fin rays (8 vs. 10–11 in O. barna,
O. dimorphicus and O. ngawa,
10 in O. chatricensis and O.
tileo, and 9 in O. dogarsinghi).
A new species of Barilius
adds to our understanding of the diversity of freshwater fishes of the Torsa River and the eastern Himalayan ecoregion. New
discoveries such as this also shows that our understanding of diversity and
conservation of freshwater fishes of this region needs to be improved and more
exploratory surveys are required.
Comparative material
Barilius barila: ZSI 54500, 85.31mm SL, Jammu, India; ZSI
F2549/2, 51.71mm SL, Belsari
River (tributary of Brahmaputra River), Assam, India; ZSI F4307/2, 51.79–84.80
mm SL, Barak River, Karong, Manipur, India; MUMF
5049, 5051, 83.22–89.53 mm SL, Khuga River, Churchandpur, Manipur, India.
Barilius vagra: MUMF Uncat, 41.67–55.69 mm
SL, Arunachal Pradesh, India.
Opsarius arunachalensis: APFS/ZSI/P-502, P-503, 110–140 mm SL, Agari River mouth, Pasighat, East
Siang District, Arunachal Pradesh, India.
Opsarius barna: MUMF 27061–27064, 73.1–83.21 mm SL, Dikrong River, Arunachal Pradesh, India.
Opsarius bendelisis: ZSI FF1357, 79.71mm SL, Cauvery River, India; ZSI
FF4269, 120mm SL, Torsa River, Cooch Behar, northern
Bengal, India; ZSI FF4270, 37.21–78.34 mm SL, Lataguri,
Jalpaiguri District, northern Bengal, India.
Opsarius chatricensis: MUMF 503/1 (holotype), 86.43mm SL, Chatrickong River, 150km from Imphal,
Ukhrul District, Manipur, India. Additional data from
Selim & Vishwanath (2002).
Opsarius dimorphicus: Data from Tilak & Husain (1990).
Opsarius dogarsinghi: MUMF 207–210, 52.89–72.26 mm SL, Chakpi
Stream, Manipur, India.
Opsarius howesi: ZSI FF2235, FF2236 61–70 mm SL, Jalpaiguri
District, northern Bengal, India.
Opsarius lairokensis: MUMF 27075, 108.45mm SL, Moreh
Bazar, Moreh, Chandel
District, Manipur, India.
Opsarius ngawa: MUMF 149 (holotype), 96.56mm SL, Sherou
River (tributary of Manipur river), 83km south of Imphal,
Manipur, India; MUMF 27056–27058, 80.1–82.96 mm SL, Singda,
Manipur, India.
Opsarius profundus: MUMF 27001 (holotype), 71.21mm SL, Koladyne River, Kolchaw, Lawntlai District, Mizoram, India.
Opsarius radiolatus: Data from Gunther (1868) and Nath et al. (2010).
Opsarius shacra: ZSI F12269, 51.15mm SL, Tribeni,
Nepal; ZSI F13405/1, 38.78–73.19 mm SL, Teesta River drainage, Kalimpong Duars & Siliguri Terai, West Bengal, India; CIFRI F10001, 72.53–73.91 mm SL;
Siang River, Pasighat, East Siang District, Arunachal
Pradesh, India.
Opsarius tileo: MUMF 27076, 128.16mm SL, Umtrao
River, Byrnihat, Norbong, Ribhoi District, Meghalaya, India.
Table 1. Morphometric data of Barilius
torsai sp. nov.
|
Parameter |
Holotype |
Paratype(s) |
|
Standard
length (SL; mm) |
71.41 |
71.46–74.56 |
|
%
SL |
|
|
|
Body depth
at dorsal-fin origin |
23.89 |
22.17–23.61 |
|
Body depth
at anal-fin origin |
17.74 |
17.79–18.24 |
|
Head length
(HL) |
26.58 |
26.66–26.84 |
|
Caudal-peduncle
length |
18.34 |
15.77–18.24 |
|
Caudal-peduncle
depth |
10.05 |
10.12–10.68 |
|
Pre-dorsal
length |
55.16 |
55.19–55.30 |
|
Pre-pelvic
length |
48.34 |
47.25–47.98 |
|
Pre-anus
length |
68.97 |
64.67–68.06 |
|
Pre-anal
length |
70.55 |
66.87–70.47 |
|
Dorsal-fin
base length |
13.98 |
13.99–14.48 |
|
Anal-fin
base length |
12.03 |
12.33–13.41 |
|
Dorsal-fin
height |
18.75 |
18.78–19.54 |
|
Pelvic-fin
length |
12.81 |
13.00–13.29 |
|
Anal fin
height |
14.33 |
14.30–14.31 |
|
Pectoral-fin
length |
18.13 |
17.18–18.10 |
|
Caudal-fin
length |
23.02 |
23.43–24.14 |
|
Body width
at anal-fin origin |
9.94 |
9.54–9.89 |
|
Body width
at dorsal-fin origin |
11.47 |
11.47–12.82 |
|
%
HL |
|
|
|
Snout
length |
29.19 |
23.64–28.69 |
|
Interorbital
width |
31.51 |
29.69–31.03 |
|
Eye orbit
diameter |
23.5 |
23.78–25.59 |
|
Mouth gape
width |
20.02 |
21.11–28.04 |
|
Head-depth
at eye |
56.95 |
58.13–66.27 |
|
Head-depth
at nape |
67.12 |
68.96–73.06 |
|
Maximum
head width |
47.26 |
47.24–47.36 |
|
Head-width
at eye |
44.89 |
44.86–44.91 |
|
Maxillary barbel length |
22.23 |
22.65–25.94 |
|
Rostral barbel length |
23.97 |
23.99–29.74 |
Table 2. Comparative measurements of body depth,
vertical bars on body and meristic counts of Barilius
torsai sp. nov. with
related species of Barilius and Opsarius.
|
Species |
Body
depth (%
SL; mm) |
Barbels
(in pair) |
Lateral
line scales |
Pre-dorsal
scales |
Vertical
bars on body |
Anal
fin branched rays |
Pectoral
fin branched rays |
|
B. torsai |
22.17–23.89 |
2 |
52–53 |
29 |
9–11 |
8 |
11 |
|
B. vagra |
25.22–26.71 |
2 |
40–45 |
21–22 |
10–14 |
10–12 |
14-15 |
|
B. barila |
23.42–24.1 |
2 |
40–45 |
22 |
14–15 |
10–11 |
12 |
|
O. shacra |
22.24–23.23 |
2 |
59–70 |
22–25 |
12–13 |
8 |
14 |
|
O. bendelisis |
22.12–26.13 |
2 |
39–46 |
18–20 |
8–12 |
7–8 |
14 |
|
O. profundus |
32.1–37.4 |
2 |
30–35 |
17–18 |
7–10 |
10 1/2 |
11 |
|
O. howesi |
30.03–31.73 |
2 |
42–43 |
17–18 |
14–15 |
7–8 |
13 |
|
O. radiolatus |
- |
2 |
56–62 |
24–25 |
Absent |
10–11 |
16 |
|
O. lairokensis |
25.54 |
2 |
41–44 |
21 |
14–16 |
11 |
13 |
|
O. arunachalensis |
20.21–20.83 |
0 |
40-45 |
21 |
Absent |
8 |
11 |
|
O. barna |
29.0–30.8 |
0 |
36–42 |
15–16 |
9–11 |
10–11 |
12 |
|
O. chatricensis |
23.2 |
0 |
38 |
15 |
7–8 |
10 |
11 |
|
O. dimorphicus |
24.9–28.8 |
1 |
60–66 |
25–27 |
Sopts |
10–11 |
12–13 |
|
O. dogarsinghi |
24.8–30.0 |
2 |
37–40 |
20 |
8–9 |
9 |
12 |
|
O. ngawa |
24.8–28.3 |
2 |
40–43 |
21–22 |
13–14 |
10–11 |
12–13 |
|
O. tileo |
29.9 |
1 |
59–4 |
28 |
Spots |
10 |
13 |
Supplementary 1. Biometric data of Barilius
torsai sp. nov.
|
Measurements |
Holotype |
Paratype
(s) |
|||||||||
|
ZSI
FF5542 |
ZSI
FF5543 |
CIFRI
F10003 |
CIFRI
F10004 |
CIFRI
F10005 |
CIFRI
F10006 |
CIFRI
F10007 |
CIFRI
F10008 |
CIFRI
F10009 |
CIFRI
F10010 |
CIFRI
F10011 |
|
|
Standard
length (mm) |
71.41 |
74.56 |
71.46 |
71.67 |
71.89 |
72.15 |
72.17 |
72.98 |
73.17 |
74.23 |
71.46 |
|
Body depth
at dorsal-fin origin |
17.06 |
16.53 |
16.07 |
16.92 |
16.95 |
17.03 |
16.6 |
16.59 |
16.58 |
16.59 |
16.04 |
|
Body depth
at anal-fin origin |
12.67 |
13.6 |
12.71 |
12.91 |
12.81 |
12.89 |
12.97 |
12.99 |
13.1 |
13.4 |
12.98 |
|
Head length
(HL) |
18.98 |
20.01 |
19.1 |
19.17 |
19.21 |
19.24 |
19.31 |
19.51 |
19.57 |
19.89 |
19.18 |
|
Caudal-peduncle
length |
13.1 |
11.76 |
12.96 |
12.91 |
13.07 |
12.46 |
12.49 |
12.79 |
12.87 |
13.54 |
12.89 |
|
Caudal-peduncle
depth |
7.18 |
7.96 |
7.62 |
7.39 |
7.28 |
7.33 |
7.34 |
7.57 |
7.66 |
7.56 |
7.26 |
|
Pre-dorsal
length |
39.39 |
41.23 |
39.48 |
39.57 |
39.73 |
39.87 |
39.88 |
40.33 |
40.41 |
41.02 |
39.44 |
|
Pre-pelvic
length |
34.52 |
35.23 |
34.12 |
34.17 |
34.12 |
34.56 |
34.19 |
34.65 |
34.67 |
35.17 |
34.29 |
|
Pre-anus
length |
49.25 |
48.22 |
48.23 |
48.27 |
48.93 |
49.11 |
48.63 |
48.72 |
48.57 |
49.19 |
48.64 |
|
Pre-anal
length |
50.38 |
49.86 |
49.89 |
49.79 |
49.32 |
50.39 |
49.89 |
49.87 |
49.98 |
50.84 |
50.36 |
|
Dorsal-fin
base length |
9.98 |
10.8 |
10.01 |
10.17 |
10.06 |
10.2 |
10.43 |
10.31 |
10.27 |
10.53 |
10.02 |
|
Anal-fin
base length |
8.59 |
10 |
8.91 |
8.97 |
8.87 |
9.27 |
9.37 |
9.67 |
9.81 |
9.92 |
8.95 |
|
Dorsal-fin
height |
13.39 |
14.57 |
13.57 |
13.78 |
13.5 |
13.67 |
13.58 |
13.89 |
14.2 |
14.46 |
13.58 |
|
Pelvic-fin
length |
9.15 |
9.91 |
9.29 |
9.42 |
9.47 |
9.39 |
9.43 |
9.57 |
9.71 |
9.79 |
9.34 |
|
Anal-fin
height |
10.23 |
10.67 |
10.22 |
10.25 |
10.28 |
10.32 |
10.32 |
10.44 |
10.47 |
10.62 |
10.22 |
|
Pectoral-fin
length |
12.95 |
12.81 |
12.35 |
12.81 |
13.01 |
12.84 |
12.79 |
12.91 |
12.88 |
12.87 |
12.4 |
|
Cauda- fin
length |
16.44 |
18 |
16.74 |
16.83 |
16.97 |
17.31 |
17.29 |
17.57 |
17.54 |
17.88 |
16.94 |
|
Body width
at anal-fin origin |
7.1 |
7.11 |
6.99 |
6.98 |
6.95 |
6.97 |
6.98 |
6.97 |
7.01 |
7.34 |
7 |
|
Body width
at dorsal-fin origin |
8.19 |
9.56 |
8.2 |
8.89 |
8.39 |
8.78 |
9.14 |
9.27 |
8.74 |
9.5 |
8.25 |
|
Snout
length |
5.54 |
4.73 |
4.87 |
4.77 |
4.78 |
4.79 |
4.76 |
5.6 |
4.98 |
5.53 |
5.5 |
|
Interorbital
distance |
5.98 |
5.94 |
5.78 |
5.7 |
5.96 |
5.78 |
5.89 |
5.86 |
5.91 |
5.91 |
5.82 |
|
Eye orbit
diameter |
4.46 |
5.12 |
4.68 |
4.56 |
4.72 |
4.65 |
4.9 |
4.65 |
4.98 |
4.97 |
4.9 |
|
Mouth gape
width |
3.8 |
5.61 |
4.04 |
4.21 |
4.73 |
5.17 |
5.35 |
5.28 |
5.38 |
5.47 |
4.05 |
|
Head depth
at eye |
10.81 |
13.26 |
12.31 |
12.15 |
11.17 |
12.37 |
11.23 |
11.35 |
12.31 |
11.94 |
11.24 |
|
Head depth
at nape |
12.74 |
14.62 |
13.69 |
13.52 |
13.25 |
13.42 |
14.02 |
14.13 |
13.79 |
13.72 |
13.7 |
|
Maximum
head width |
8.97 |
9.46 |
9.03 |
9.08 |
9.08 |
9.09 |
9.146 |
9.24 |
9.25 |
9.4 |
9.06 |
|
Head width
at eye |
8.52 |
8.98 |
8.57 |
8.61 |
8.62 |
8.64 |
8.67 |
8.76 |
8.78 |
8.93 |
8.61 |
|
Maxillary barbel length |
4.22 |
5.19 |
4.33 |
4.97 |
4.69 |
4.74 |
5.01 |
5.02 |
4.98 |
5.11 |
4.35 |
|
Rostral barbel length |
4.55 |
5.95 |
4.59 |
4.6 |
4.7 |
5.12 |
5.32 |
4.79 |
4.89 |
5.39 |
4.62 |
For figure
& images - - click here
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