Journal of Threatened
Taxa | www.threatenedtaxa.org | 26 February 2026 | 18(2): 28329–28342
ISSN 0974-7907 (Online) | ISSN 0974-7893 (Print)
https://doi.org/10.11609/jott.10272.18.2.28329-28342
#10272 | Received 24 November 2025 | Final received 01 February 2026 |
Finally accepted 05 February 2026
Local knowledge, attitudes, and
perceptions of ecosystem services and disservices provided by the Painted Stork
Mycteria leucocephala
Pennant, 1769 (Aves: Ciconiidae) in
northern India: insights for conservation
Yashmita-Ulman 1 & Manoj Singh 2
1 Department of Silviculture &
Agroforestry, College of Horticulture & Forestry, Acharya Narendra Deva
University of Agriculture and Technology, Kumarganj, Ayodhya, Uttar Pradesh 224229, India.
2 Department of Zoology, Kalinga
University, Naya Raipur, Chhattisgarh 492012, India.
1 yashmita-agf@nduat.org, 2
manoj.singh@kalingauniversity.ac.in (corresponding author)
Editor: H. Byju,
Coimbatore, Tamil Nadu, India. Date of publication: 26 February 2026 (online & print)
Citation: Yashmita-Ulman & M. Singh (2026). Local
knowledge, attitudes, and perceptions of ecosystem services and disservices
provided by the Painted Stork Mycteria leucocephala Pennant, 1769 (Aves: Ciconiidae)
in northern India: insights for conservation. Journal of Threatened Taxa 18(2): 28329–28342. https://doi.org/10.11609/jott.10272.18.2.28329-28342
Copyright: © Yashmita-Ulman & Singh 2026. Creative Commons Attribution 4.0
International License. JoTT allows unrestricted use,
reproduction, and distribution of this article in any medium by providing
adequate credit to the author(s) and the source of publication.
Funding: This study was carried out through funding provided by the Wildlife Trust of India and Fondation Segre.
Competing interests: The authors declare no competing interests.
Author details:
Dr. Yashmita-Ulman, assistant professor in the Department of Silviculture & Agroforestry, ANDUAT, Ayodhya, Uttar Pradesh. She specialises in the field areas of agroforestry and wildlife conservation and management. Dr. Manoj Singh, assistant professor in the Department of Zoology, Kalinga University, Naya Raipur, Chhattisgarh. He is specialized in bird acoustics and wildlife conservation.
Author contributions: Y-U conceived and designed the study, conducted fieldwork, and wrote the final draft of the manuscript. MS performed the field work, compiled the data and analyzed the data. Both authors read and approved the final manuscript.
Acknowledgements: We thank the funding agencies, Wildlife Trust of India and Fondation Segre, and the villagers for their permission and constant support in conducting this survey.
Abstract: Understanding the knowledge,
attitudes and perceptions of the local communities towards the target species
before implementing any awareness-based programmes is
essential for the success and long-term protection of the species. No studies have yet been conducted on local
knowledge, attitudes, perceptions of ecosystem services and disservices,
threats and possible recommendations for conservation of the Painted Stork Mycteria leucocephala in
northern India. To fill this gap, a study was conducted in August–September
2024 in villages of Etawah District, Uttar Pradesh,
India, where Painted Stork builds nests in farmland in human-dominated areas.
The study employed a questionnaire survey, containing both closed and
open-ended questions, administered to 200 respondents. All the respondents
identified the species, and a majority knew its local name, breeding ecology,
foraging ecology, and population trend. A majority of the people (52%) liked
the species, supported conservation of the species (51%), felt the need for
awareness programmes (52%), and were ready to
participate in nest monitoring training (47%) & awareness programmes (58%). At the same time, a majority of the people
were reluctant to report hunting (48%), reluctant to take the injured birds to
veterinary doctors (46%), and lacked knowledge on the
bird’s conservation status (64%). People perceived 12 ecosystem services and
five ecosystem disservices provided by Painted Stork, with pest control (50%)
being the highest cited service and crop destruction (72%) the most cited
disservice. According to the respondents, the highest threat to the species is
wetland loss (59%). A majority recommended that conservation efforts should
focus on organizing awareness programmes (52%) and
implementing wetland management and conservation (50%). Furthermore,
respondents believed the community should take the initiative in these efforts
(48%). This study emphasizes intensified awareness programs, wetland
conservation and management and protection of nesting trees, among other
measures, to ensure the conservation of this species outside protected areas.
It provides the essential baseline for conservationists, researchers, and
policy makers to design effective conservation and management strategies
involving local communities and policies for research and conservation of
Painted Stork.
Keywords: Etawah,
Farhadpura, human-dominated landscapes, nesting
trees, questionnaire surveys, recommendations, socio-demography
characteristics, threats, Usrai, Uttar Pradesh.
INTRODUCTION
Anthropogenic
factors such as deforestation, urbanization, encroachment, human-wildlife
negative interactions have driven the species to the brink of extinction and
led to wildlife population declines (Taylor-Brown et al. 2019; Lewis et al.
2021; Lees et al. 2022). The scientific community has tried to invent many
different possibilities, such as creating wildlife sanctuaries (Gorbunov et al. 2019; Volenec
& Dobson 2020), restoration of degraded areas (Possingham
et al. 2015; Volis 2019), and ex situ conservation (Canessa et al. 2015). But focusing on these techniques is
not sufficient, as many of the threatened species are found in human-dominated
landscapes (Bracebridge et al. 2013; Yashmita-Ulman et al. 2018; 2021a,b;
Ceballos et al. 2019). The co-existence of people with wildlife is ultimately
seen as a necessity to achieve wildlife conservation (Chakanyuka
& Utete 2022), emphasizing the importance of the
community’s ecological knowledge and attitudes towards wildlife conservation
(Gupta et al. 2023; Manigandan et al. 2024).
As locals
interact with native species routinely, they have a good insight
(Romero-Bautista et al. 2020) into the aspects of species distribution,
breeding, diet, associated problems, services and threats (Gaston et al. 2018; Kross et al. 2018) as is the case with the Painted Stork Mycteria leucocephala,
which is commonly seen foraging and building nesting colonies in the wetlands (Yashmita-Ulman 2022, 2023; Yashmita-Ulman
& Singh 2022, 2025) and agricultural fields (Yashmita-Ulman
& Singh 2021) in Uttar Pradesh. Understanding the local knowledge and
involving the communities in the decision-making process can make them actively
participate in conservation programmes. This can lead
to the effective design of species conservation management strategies (Katuwal et al. 2021; Messina et al. 2023). Benefit sharing
related to conservation-related activities can inculcate positive attitudes
towards the species (Bajracharya et al. 2007),
whereas events such as crop-raiding, livestock depredation, which are the costs
related to conservation, develop negative attitudes towards the species (Mehta
& Heinen 2001; Walpole & Goodwin 2001; Yashmita-Ulman
et al. 2020). Evidence shows that communities hold negative attitudes towards
birds such as Sarus Crane (Nevard
et al. 2019), raptors (Dabone et al. 2022),
Black-necked Crane (Zhong et al. 2023), for crop damage, making noise (Leong et
al. 2020), and spreading disease (Green & Elmberg
2014). Certain communities also appreciate birds for their ecosystem services,
such as decorative value (Baya Weaver, Yashmita-Ulman et al. 2017), scavenging (vultures,
Morales-Reyes et al. 2018), pest-control (owls, Iniguez-Gallardo et al. 2024),
aesthetic value (Sarus crane, Bhattarai et al. 2025),
and revere them for their cultural values (Hooded vultures, Dabone
et al. 2022). Therefore, the attitudes of the community towards the species
need to be understood as these play an important role in the acceptance or
rejection of conservation-related activities and management practices (Winter
et al. 2005; Byju et al. 2024). The attitude of
people towards a species is influenced by factors such as age, caste, gender,
size of landholding, education, religion, and distance from forests
(Arjunan
et al. 2006; Ntuli et al. 2019). These attitudes can
also be changed through organizing effective conservation programmes
based on the prior understanding of the knowledge and attitudes of the
community towards the species. For effective implementation of any
community-based conservation activity, the first step is to understand the
local knowledge, perceptions and attitudes of the local people towards the
species in question.
The Painted
Stork, locally known as Dhek, is an ibis-like stork
with a long, tapering and drooping bill, 93–102 cm in length, and it weighs
around 2–3.5 kg. It is distributed across southwestern Pakistan, India, and Sri
Lanka, and into Indochina and southern China. It is found in shallow freshwater
ecosystems such as lakes, marshlands, rivers, paddy fields, and wetlands
(Cornell Lab of Ornithology 2025). It is a locally migratory species in Uttar
Pradesh, and its breeding season lasts from August/September to February. The
global population was estimated to be around 25,000 birds in 1994 (Cornell Lab
of Ornithology 2025). The bird species has been upgraded to Schedule II, Part B
of the Wildlife (Protection) Amendment Act, 2022 (WPAA 2022) from Schedule IV
of the Wildlife (Protection) Act, 1972 (WPA 1972). Though the IUCN status of
this bird is ‘Least Concern’ (IUCN 2025), it locally faces the threat of
hunting, habitat loss, and degradation (Cornell Lab of Ornithology 2025). This
species is mainly hunted for its meat and eggs (Cornell Lab of Ornithology
2025) as it forms a cheap source of protein for the local communities. In Uttar
Pradesh, the Painted Stork prefers wetlands and agricultural fields for nesting
(Tiwary et al. 2014) and foraging, which are present
in human-dominated landscapes, not under the direct control of the forest
department but under the community governance, increasing the chances of
hunting and habitat destruction that go unreported. The most effective way to
protect this species and its nesting sites is to integrate its conservation and
management with local communities and their livelihoods.
Many
studies have been conducted in different countries across the world that
document the local knowledge and attitudes of people towards bird species
conservation (Jacobson et al. 2003; Mmassy & Roskaft 2013; Cortes-Avizanda et al. 2018; Gaston et al.
2018; Kross et al. 2018; Katuwal
et al. 2021, 2024; Zhong et al. 2023). In India, a handful of such studies
exist on wildlife (Mir et al. 2015; Talukdar & Gupta 2018; Karanth et al. 2019; Gupta et al. 2023; Bhaskaran
& Nilon 2025), and birds in particular (Acharya
et al. 2009; Aiyadurai & Banerjee 2020; Aiyadurai et al. 2023; Tiwari et al. 2023; Das et al.
2025), but there are none specifically on Painted Stork. Considering the
knowledge gap on the subject and the species, this study aims to assess: 1)
people’s ecological knowledge on Painted Stork, 2) their attitudes towards the
species, 3) people’s perception of threats faced by the species, 4) perceptions
on ecological services and disservices provided by the Painted Stork, and 5)
possible recommendations to conserve the species as suggested by the community.
MATERIALS
AND METHODS
Study area
The field
sites for this study are centred around the villages
of Farhadpura (26.888° N & 79.161° E) and Usrai (Usrahar) (26.919° N &
79.034° E) (Figure 1), both situated within the Barhpura
Block of the Etawah District in southwestern Uttar
Pradesh. These localities are part of the Ganga-Yamuna Doab—a region defined by
its position between the two major rivers, the Yamuna and its main tributary,
the Chambal (KVK 2025). The district covers an area of approximately 2,311 km²,
with the district headquarters in Etawah Town (GoUP 2025). The district experiences a sub-humid climate,
with an average annual rainfall of about 792 mm, most of which falls between
June and September during the southwest monsoon (KVK 2025). Temperatures peak
in May, reaching daily maxima averaging up to 42 °C. Much of the area is
agricultural, dominated by wheat, rice, and sugarcane crops (KVK 2025), but
remnant patches of natural vegetation—especially along water bodies, agricultural
fields and communal/village lands—serve as critical nesting sites for waterbirds. The Sarsai Nawar Wetland, recognized as a Ramsar
site, lies in proximity and underlines the conservation importance of this
region for waterbirds.
The Painted
Stork is a colonial nester. In both the villages, these birds build nests on
the trees of Terminalia arjuna and Ficus
religiosa found in agricultural fields (Image 1).
Method
The
questionnaire surveys were carried out for two months, August–September 2024.
Before starting with the formal data collection, an introductory informal group
discussion was conducted with the residents of both the villages with the help
of the village headman or gaon pradhan.
Then, a preliminary pilot survey was carried out by interviewing 40
respondents, based on which a few improvements were made in the questionnaires.
The respondents were also asked to list the benefits and problems caused by the
Painted Stork. Based on these answers, a list of 12 ecosystem services and five
ecosystem disservices were included in the final questionnaires. The ecosystem
services were further classified into three sections, namely, regulation &
maintenance, cultural, and provisioning services (Zhong et al. 2023). The
improvised questionnaire had a set of closed and open-ended questions (See
supplementary file S1) administered to 100 randomly selected respondents (as
the nesting trees were in or around agricultural fields) from each village,
making a total of 200 respondents. The questions asked were pertaining to 1)
knowledge on Painted Stork (species identification, local name, habitat,
nesting trees, breeding season, foraging locations, diet, population
fluctuations over the years, conservation status of species), 2) attitudes
towards Painted Stork (thought of it as a harmful species, whether they liked
the species, supported its conservation, whether they cut its nesting trees,
opinions on need for awareness programmes,
willingness to participate in nest monitoring training programmes
and awareness programmes, report hunting, take
injured birds to veterinary doctor), 3) threats faced as perceived by people,
4) recommendations to conserve the species as suggested by people, and 5)
perceptions of ecosystem services and disservices provided by the species (respondents
were asked to choose between the 12 ecosystem services namely pest control,
eats snakes and nutrient cycling under regulation and maintenance ecosystem
service, medicinal value and meat value under provisioning ecosystem service,
sense of pride, existence value, aesthetic value, conservation value,
conservation flagship, educational value and ecotourism under cultural
ecosystem service and five ecosystem disservices namely destroys crops, eat
fishes, makes noise, makes area dirty and spreads diseases). To establish
whether the participants could identify the species and its local name, a
photo-elicitation approach was used during the surveys. Before the start of
every interview, the verbal consent of the respondents was taken, and they were
informed that the data would be used only for research purposes. The interviews
were conducted in the local Hindi language using simple local terms. Each
interview took approximately 40 minutes to complete. For data analysis, the
respondent’s knowledge about the species was evaluated by marking one point for
each correctly answered question and zero points for each wrong answer.
Results and Discussion
Socio-demographic
characteristics of respondents
Out of the
200 respondents, 66% (n = 132) were male, and 34% (n = 68) were female (Table
1). The respondents’ ages ranged from 16–85 years, with the majority falling
within the age range of 25–54 (Table 1). The majority of the respondents were
engaged in agriculture and received secondary level education till 12th
class (Table 1).
Respondent’s
ecological knowledge about the species
All the
respondents were able to identify the Painted Stork, and more than half knew
its local name (Figure 2). While most respondents could not correctly identify
the bird’s habitats, they were knowledgeable about the breeding season and
preferred nesting trees of the species. The respondents mentioned that the
Painted Storks arrive during the Diwali season (i.e., September/October) and
avoid nesting on sacred Peepal Ficus
religiosa trees used for prayer. Most of the
respondents knew that wetlands and agricultural lands were the species’ most
preferred foraging sites, and that the species fed on fish, snakes,
crustaceans, snails, shrimps, crabs, insects, and frogs. The familiarity of the
local people with this species might be because the Painted Stork has an
attractive appearance, a huge body and builds nests in and around farmlands and
wetlands, close to human habitations. Similarly, many other studies have
reported that people possess good knowledge of birds around them (Mmassy & Roskaft 2013; Katuwal et al. 2024; Das et al. 2025; Nima
et al. 2025). The findings also report declines in the population of species
(Figure 2), which might be due to the comparison with their past experiences.
Interestingly, the population trend of this species shows that its population
is increasing (BirdLife International 2025),
suggesting a local decline in the species population. The majority of the
people were unaware of the conservation status of the species and did not know
its habitat (Figure 2), which also shows the knowledge gap. Some studies have
reported that educated people have greater awareness of the conservation status
of birds and their habitat (Ortega-Lasuen et al.
2023; Katuwal et al. 2024).
Respondent’s
attitudes towards the species
Amongst the
respondents, 44% (n = 87) perceived the Painted Stork as a harmful species
(Figure 3), primarily because its excreta fell on and destroyed crops. The
majority of villagers supported species conservation and cherished the birds
(Figure 3), as they were accustomed to living with them like family members and
appreciated their elegant appearance. Furthermore, the respondents agreed that
awareness programmes were necessary and expressed
readiness to participate in nest monitoring and awareness programmes
(Figure 3), consistent with findings from a study on the Sarus
Crane in Nepal (Bhattarai et al. 2025). Nearly an equal number of people agreed
and disagreed about taking injured birds to the veterinarians, with many
refusing to report hunting (Figure 3). This reluctance likely stems from a
desire to avoid legal repercussions. Consequently, hunting in villages remains
underreported, causing this threat to rise slowly.
Perceived
threats to the species
According
to the opinion of the respondents, the highest threat faced by the Painted
Stork is wetland loss and hunting (Figure 4). Similar threats have been
reported by Herzon & Mikk
(2007), Stanton et al. (2018) and Katuwal et al.
(2021). Villagers confirmed that most wetlands were encroached upon for
agricultural expansion and that this species was hunted for meat. During our
field visits, some people were seen pelting stones at the chicks in the nest.
There are reports that confirm this species is hunted for food (BirdLife International 2025). Although we observed one
instance where a tree was cut down to avoid nesting, while a few villagers
confirmed this during surveys, the majority denied it.
Perceived
ecosystem services and disservices
In all, the
respondents recognised 12 ecosystem services (three
regulation and maintenance ecosystem services, two provisioning ecosystem
service and seven cultural ecosystem services) and five ecosystem disservices
(Figures 5 & 6). The most-cited regulation and maintenance ecosystem
service was pest control; the most-cited provisioning ecosystem service was
meat value, and the most-cited cultural ecosystem service was ecotourism
(Figure 5). Because the diet of the Painted Stork consists of agricultural
pests – including insects, snails (Ali & Ripley 1987; Parasharya
& Naik 1990; Sridhar et al. 2002; Urfi 2002), and
snakes (Urfi 1988) – humans might appreciate this
bird’s role as a natural predator. The positive attitude towards this species
might also have been developed as people acknowledged the ecosystem services
provided by the Painted Stork (Whelan et al. 2008; Mariyappan
et al. 2023). The most-cited ecosystem disservices were crop destruction and
fish predation (Figure 6). Bird excreta that drop on nesting trees and crops
below (observed by Yashmita-Ulman) form a white layer
on leaves, disrupting photosynthesis and reducing crop yield. Since the species
is primarily piscivorous (Urfi 2011), this predation
on fish—a food source for local people—results in human-wildlife negative
interactions. The number of services and disservices mentioned by each
respondent ranged 1–3, with the majority citing one ecosystem service in each
category (Figures 7, 8, & 9) and two ecosystem disservices (Figure 10).
Conservation
implications
The
majority of respondents suggested conservation measures such as organizing
awareness programs, wetland management and conservation, establishment of
protected areas, and hunting control (Figure 11). The respondents strongly
believed that the community should take the initiative for species
conservation, consistent with findings from the study on Sarus
Crane in Nepal (Bhattarai et al. 2025).
CONCLUSION
The
ecological knowledge, attitudes, and conservation recommendations of the local
people revealed through this study increase the chances of positive impacts
from awareness activities in this area. This positive mindset is necessary for
species conservation in human-dominated landscapes. The government, along with
the non-governmental organizations working in this region, must take immediate
steps to spread awareness and provide incentives to the people involved in the
conservation of this species. Policies must be framed such that people are
encouraged to protect and plant the preferred nesting trees of the Painted
Stork, such as Ficus religiosa,
Syzygium cumini, and
Terminalia arjuna report hunting activities and manage wetlands
sustainably. This study provides the first-ever insight into people’s
knowledge, attitudes, perceptions of ecosystem services and disservices
provided by the Painted Stork from northern India. This study provides the
baseline data for further field-based research aimed at understanding the
interaction dynamics between birds and humans outside protected areas.
Table 1. Socio-demography
of the respondents
who participated in the interviews.
|
Variables |
Categories |
Number (N) |
Percentage (%) |
|
Gender |
Male |
132 |
66 |
|
|
Female |
68 |
34 |
|
Age |
Early working age (16–24) |
37 |
18 |
|
|
Prime working age (25–54) |
78 |
39 |
|
|
Mature working age (55–64) |
61 |
31 |
|
|
Elderly age group (> 65) |
24 |
12 |
|
Educational Attainment |
Illiterate (no formal
education) |
29 |
15 |
|
|
Primary level (< 8 class) |
46 |
23 |
|
|
Secondary level (9–12 class) |
74 |
37 |
|
|
Bachelor & above |
51 |
25 |
|
Occupation |
Agriculture |
113 |
57 |
|
|
Business |
34 |
17 |
|
|
Government job |
20 |
10 |
|
|
Private job |
29 |
14 |
|
|
Others |
4 |
2 |
For
figures, images, Appendix - - click here for full PDF
REFERENCES
Acharya, B.K., B. Chettri & L. Vijayan (2009). Indigenous
knowledge of Lepcha community for monitoring and
conservation of birds. Indian Journal of Traditional Knowledge 8(1):
65–69.
Aiyadurai, A. &
S. Banerjee (2020). Bird conservation from obscurity to popularity:
a case study of two bird species from Northeast India. GeoJournal
85: 901–912. https://doi.org/10.1007/s10708-019-09999-9
Aiyadurai, A., S.
Banerjee, Y. Patil & S. Joshi (2023). Human-bird
relations in India — An interdisciplinary Study. Indian
Institute of Technology, Gandhinagar and Environmental Humanities Research
Group, 55 pp.
Ali, S. & S.D. Ripley (1987). Compact
Handbook of the Birds of India and Pakistan. Oxford University Press,
Delhi, India, 737 pp.
Arjunan, M., C.
Holmes, J-P. Puyravaud & P. Davidar
(2006). Do developmental initiatives influence local attitudes toward
conservation? A case study from Kalakad-Mundanthurai
Tiger Reserve, India. Journal of Environmental Management 79: 188–197. https://doi.org/10.1016/j.jenvman.2005.06.007
Bajracharya, S., G.B. Gurung & K. Basnet (2007). Learning from community participation in conservation
area management. Journal of Forest and Livelihood 6: 54–66. https://www.nepjol.info/index.php/JFL/article/view/2325
Bhaskaran, V. & C.H.
Nilon (2025). Exploring
perceptions and attitudes towards wildlife in urban home gardens in Bangaluru, India. Urban Ecosystems 28: 107. https://doi.org/10.1007/s11252-025-01723-5
Bhattarai, B.P., H.B. Katuwal, S. Regmi, A. Nepali, R.N. Suwal, R.
Acharya, K.C. Sabin, B. Aryal, K. Tamang, B. Rawal,
A. Basnet, B.D. Baral, S. Devkota,
S. Parajuli, N. Regmi, P.
Kandel, B. Subedi, H.S. Giri,
G.S. Gurung, J.L. Belant & H.P. Sharma (2025). Knowledge,
attitudes, and conservation threats to globally vulnerable Sarus
cranes in Lumbini province, Nepal. Discover Conservation 2: 15. https://doi.org/10.1007/s44353-025-00034-2
BirdLife
International (2025). Species factsheet: Mycteria
leucocephala. https://datazone.birdlife.org/species/factsheet/painted-stork-mycteria-leucocephala.
Accessed on 15.xi.2025.
Bracebridge, C., T.R.B.
Davenport, V.F. Mbofu & S.J. Marsden (2013). Is There a
Role for Human-Dominated Landscapes in the Long-Term Conservation Management of
the Critically Endangered Kipunji (Rungwecebus
kipunji)? International Journal of Primatology 34:
1122–1136. https://doi.org/10.1007/s10764-013-9719-3
Byju, H., H. Maitreyi, S. Ravichandran & N. Raveendran (2024). Avifaunal
diversity and conservation significance of coastal ecosystems on Rameswaram
Island, Tamil Nadu, India. Journal of Threatened Taxa 16(12):
26198–26212. https://doi.org/10.11609/jott.9248.16.12.26198-26212
Canessa, S., S.J.
Converse, M. West, N. Clemann, G. Gillespie, M. McFadden,
A.J. Silla, K.M. Parris & M.A. McCarthy (2015). Planning
for ex-situ conservation in the face of uncertainty. Conservation Biology
30(3): 599–609. https://doi.org/10.1111/cobi.12613
Ceballos, G., P.R. Ehrlich, J. Pacheco, N. Valverde-Zuniga & G.C. Daily
(2019). Conservation in human-dominated landscapes: Lessons from the
distribution of the Central American squirrel monkey. Biological
Conservation 237: 41-49. https://doi.org/10.1016/j.biocon.2019.06.008
Chakanyuka, T. &
B. Utete (2022). Adaptive
co-management, co-existence or just wildlife conservation? Case study of the
human and Nile crocodile Crocodylus miloticus conflicts in Ngezi
Dam, Mashonaland West, Zimbabwe. African Journal of Ecology 60: 759–768.
https://doi.org/10.1111/aje.12974
Cornell Lab of Ornithology (2025). Cornell Lab
of Ornithology, Birds of the World. https://birdsoftheworld.org/bow/species/paisto1/cur/introduction#distrib.
Accessed on 17.xi.2025.
Cortes-Avizanda, A., B. Martin-Lopez, O. Ceballos & H.M. Pereira
(2018). Stakeholders perceptions of the Endangered Egyptian vulture: insights
for conservation. Biological Conservation 218: 173–180. https://doi.org/10.1016/j.biocon.2017.09.028
Dabone, C., A. Oueda, L.J. Thompson, J.B. Adjakpa
& P.D.M. Weesie (2022). Local
perceptions and sociocultural value of Hooded Vultures Necrosyrtes
monachus in Burkina Faso, West Africa. Ostrich 93(4): 233–247. https://doi.org/10.2989/00306525.2022.2120558
Das, S., A. Srivastava & U. Hore (2025). Farmer’s
knowledge on bird conservation in the agricultural landscape of Gangetic plain.
Ornithology Research 33: 26. https://doi.org/10.1007/s43388-025-00231-6
Gaston, K.J., D.T.C. Cox, S.B. Canavelli, D.
Garcia, B. Hughes, B. Maas, D. Martinez, D. Ogada
& R. Inger (2018). Population abundance and ecosystem service
provision: the case of birds. Bioscience 68: 264–272. https://doi.org/10.1093/biosci/biy005
Gorbunov, M.A., A.V.
Fadeeva, V.B. Shirshikov,
P.A. Matveev, O.V. Popova, M.Y. Mitrofanova,
J.Y. Bakaeva & N.A. Mashkin
(2019). Nature protection potential of wildlife sanctuary: protection and
preservation of its ecological biodiversity. Ekoloji
28(107): 5033–5037.
GoUP (2025). Government of Uttar Pradesh. https://etawah.nic.in/about-district/
Accessed on 10.xi.2025.
Green, A.J. & J. Elmberg (2014). Ecosystem
services provided by waterbirds. Biological
Reviews 89: 105–122. https://doi.org/10.1111/brv.12045
Gupta, T., E.J. Milner-Gulland, A. Dias & D. Karnad
(2023). Drawing on local knowledge and attitudes for the conservation of
critically endangered rhino rays in Goa, India. People and Nature 5:
645–659. https://doi.org/10.1002/pan3.10429
Herzon, I. &
M. Mikk (2007). Farmers’
perceptions of biodiversity and their willingness to enhance it through agri-environment schemes: a comparative study from Estonia
and Finland. Journal for Nature Conservation 15: 10–25. https://doi.org/10.1016/j.jnc.2006.08.001
Iniguez-Gallardo, V., F. Reyes-Bueno, I. Gonzalez-Coronel, J. Freile & L. Ordonez-Delgado (2024).
Perceptions, knowledge, and emotions about owls in southern Ecuador. Journal
of Ethnobiology 44(2): 98–111. https://doi.org/10.1177/02780771241250129
IUCN (2025). The IUCN Red list of threatened species: search
results for ‘Birds’. International Union for Conservation of Nature. https://www.iucnredlist.org/search?query=Birds&searchType=species.
Accessed on 25.x.2025.
Jacobson, S.K., K.E. Sieving, G.A. Jones & A. Van Doorn (2003). Assessment of farmer attitudes
and behavioral intentions toward bird conservation on organic and conventional
Florida farms. Conservation Biology 17: 595–606. https://doi.org/10.1046/j.1523-1739.2003.01472.x
Karanth, K.K., S.
Jain & E. Weinthal (2019).
Human-wildlife interactions and attitudes towards wildlife and wildlife
reserves in Rajasthan, India. Oryx 53(3): 523–531. https://doi.org/10.1017/S0030605317001028
Katuwal, H.B., H.P.
Sharma, P. Rokka, N.K. Das & R-C. Quan (2024). Knowledge,
attitudes and conservation challenges for the Lesser Adjutant in Nepal. Global
Ecology and Conservation 49: e02795.
https://doi.org/10.1016/j.gecco.2023.e02795
Katuwal, H.B., M.
Zhang, H.S. Baral, H.P. Sharma & R.-C. Quan (2021). Assessment of farmers’ knowledge and perceptions towards farmland birds show the need of conservation interventions. Global
Ecology and Conservation 27: e01563. https://doi.org/10.1016/j.gecco.2021.e01563
Kross, S.M., K.P.
Ingram, R.F. Long & M.T. Niles (2018). Farmer
perceptions and behaviors related to wildlife and on-farm conservation actions.
Conservation Letters 11(1): e12364.
https://doi.org/10.1111/conl.12364
KVK (2025). Krishi Vigyan Kendra Etawah.
https://etawah.kvk4.in/district-profile.php. Accessed
on 10.xi.2025.
Lees, A.C., L. Haskell, T. Allinson,
S.B. Bezeng, I.J. Burfield,
L.M. Renjifo, K.V. Rosenberg, A. Viswanathan
& S.H.M. Butchart (2022). State of the world’s birds. Annual Review of
Environment and Resources 47(1): 231–260. https://doi.org/10.1146/annurev-environ-112420-014642
Leong, R.A.T., T.K. Fung, U. Sachidhanandam,
Z. Drillet, P.J. Edwards & D.R. Richards (2020). Use of structural
equation modelling to explore influences on perceptions
of ecosystem services and
disservices attributed to birds
in Singapore. Ecosystem Services 46: 101211. https://doi.org/10.1016/j.ecoser.2020.101211
Lewis, J.S., S. Spaulding, H. Swanson, W. Keeley, A.R. Gramza, S. VandeWoude & K.R. Crooks
(2021). Human activity influences
wildlife populations and activity
patterns: implications for
spatial and temporal refuges. Ecosphere
12(5): e03487. https://doi.org/10.1002/ecs2.3487
Manigandan, S., H. Byju & P. Kannan (2024). Harmonizing
ecology and society: an integrated analysis of vulture conservation in the Nilgiri Biosphere Reserve, India. Journal of Threatened
Taxa 16(6): 25330–25344. https://doi.org/10.11609/jott.8915.16.6.25330-25344
Mariyappan, M., M.
Rajendran, S. Velu, A.D. Johnson, G.K. Dinesh, K. Solaimuthu, M. Kaliyappan &
M. Sankar (2023). Ecological
role and ecosystem services of birds: a review. International Journal of
Environment and Climate Change 13(6): 76–87. https://doi.org/10.9734/ijecc/2023/v13i61800
Mehta, J.N. & J.T. Heinen (2001). Does
community-based conservation shape favorable attitudes among locals? An
empirical study from Nepal. Environmental Management 28: 165–177. https://doi.org/10.1007/s002670010215
Messina, T., R. Figueira & J.M.L. Santos
(2023). Integrating local and ecological knowledge to assess the benefits of
trees for ecosystem services: a holistic process-based methodology. Ecosystem
Services 63: 101556. https://doi.org/10.1016/j.ecoser.2023.101556
Mir, Z.R., A. Noor, B. Habib & G.G. Veeraswami
(2015). Attitudes of local people towards wildlife conservation: a case study
from Kashmir valley. Mountain Research and Development 35(4): 392–400. https://doi.org/10.1659/MRD-JOURNAL-D-15-00030.1
Mmassy, E.C. & E.
Roskaft (2013). Knowledge of birds
of conservation interest among the people living close
to protected areas in Serengeti, Northern
Tanzania. International Journal of Biodiversity Science, Ecosystem
Services and Management 9(2): 114–122.
https://doi.org/10.1080/21513732.2013.788566
Morales-Reyes, Z., B. Martin-Lopez, M. Moleon,
P. Mateo-Tomas, F. Botella, A. Margalida,
J.A. Donazar, G. Blanco, I. Perez & J.A.
Sanchez-Zapata (2018). Farmer perceptions of the ecosystem services
provided by scavengers: what, who and to who. Conservation Letters
11(2): 1–11. https://doi.org/10.1111/conl.12392
Nevard, T.D., I.
Leiper, G. Archibald & S.T. Garnett (2019). Farming and
cranes on the Atherton Tablelands, Australia. Pacific Conservation Biology
25: 184–192. https://doi.org/10.1071/PC18055
Nima, P., T. Dorji, M.S. Rana & T. Dorji
(2025). Knowledge, attitude, perceived threats and conservation challenges of
the critically endangered White-bellied Heron, Ardea
insignis, in Bhutan. Global Ecology and Conservation 58: e03484. https://doi.org/10.1016/j.gecco.2025.e03484
Ntuli, H., S.C. Jagers, A. Linell, M. Sjostedt & E. Muchapondwa
(2019). Factors influencing local communities’ perceptions towards conservation
of transboundary wildlife resources: the case of the Great Limpopo
Trans-frontier Conservation Area. Biodiversity and Conservation 28:
2977–3003. https://doi.org/10.1007/s10531-019-01809-5
Ortega-Lasuen, U., O. Pedrera,
E. Telletxea, O. Barrutia
& J.R. Diez (2023). Secondary students’ knowledge on birds and
attitudes towards conservation: evaluation of an environmental education
program. International Journal of Environmental Research and Public Health
20: 5769. https://doi.org/10.3390/ijerph20105769
Parasharya, B.M. &
R.M. Naik (1990). Ciconiiform birds
breeding in Bhavnagar city, Gujarat, pp. 429–445. In: Daniel, J.C. & J.C.
Serrao (eds.). Conservation in Developing Countries: Problems and Prospects.
Bombay Natural History Society and Oxford University Press, Mumbai.
Possingham, H.P., M.
Bode & C.J. Klein (2015). Optimal Conservation Outcomes Require Both
Restoration and Protection. PLoS Biology
13(1): e1002052. https://doi.org/10.1371/journal.pbio.1002052
Romero-Bautista, Y.A., A.I. Moreno-Calles, F. Alvarado-Ramos, M. Reyes
Castillo & A. Casas (2020). Environmental interactions
between people and birds in semiarid lands of the Zapotitlan
Valley, Central Mexico. Journal of Ethnobiology and Ethnomedicine 16:
32. https://doi.org/10.1186/s13002-020-00385-1
Sridhar, S., A.K. Chakravarthy, K. Srihari & N.A. Prakash (2002). Priorities
for conservation of Kokre Bellur
heronry in Karnataka, south India, pp. 123–127. In: Rahmani,
A.R. & G. Ugra (eds.). Birds of Wetlands
and Grasslands: Proceedings of the Salim Ali Centenary Seminar on Conservation
of Avifauna of Wetlands and Grasslands. Bombay Natural History Society, Mumbai.
Stanton, R.L., C.A. Morrissey & R.G. Clark (2018). Analysis of
trends and agricultural drivers of farmland bird declines in North America: a
review. Agriculture, Ecosystems & Environment 254: 244–254. https://doi.org/10.1016/j.agee.2017.11.028
Talukdar, S. & A. Gupta (2018). Attitudes
towards forest and wildlife, and conservation-oriented traditions, around Chakrashila Wildlife Sanctuary, Assam, India. Oryx
52(3): 508–518. https://doi.org/10.1017/S0030605316001307
Taylor-Brown, A., R. Booth, A. Gillett, E. Mealy, S.M. Ogbourne, A. Polkinghorne & G.C. Conroy (2019). The impact
of human activities on Australian wildlife. PLoS
ONE 14(1): e0206958. https://doi.org/10.1371/journal.pone.0206958
Tiwari, G., P. Pandey, R. Kaul & R. Singh (2023). Farmer’s
perception of the ecosystem services provided by diurnal raptors in arid
Rajasthan. PeerJ 11: e15996. https://doi.org/10.7717/peerj.15996
Tiwary, N.K., B.B.
Sharma & A.J. Urfi (2014). Two new nesting colonies of Painted
Stork Mycteria leucocephala
from northern India. Indian Birds 9(4): 85–88.
Urfi, A.J. (1988). Painted Stork Mycteria leucocephala (Pennant) swallowing
a snake. Journal of the
Bombay Natural History Society 86: 96.
Urfi, A.J.
(2002). Waders and other wetland birds on Byet Dwarka
Island, Gulf of Kutch, Western India. Wader Study Group Bulletin 99:
31–34.
Urfi, A.J.
(2011). Foraging ecology of the Painted Stork (Mycteria
leucocephala): A review. Waterbirds
34(4): 448-–456. https://doi.org/10.1675/063.034.0407
Volenec, Z.M. &
A.P. Dobson (2020). Conservation value of small reserves. Conservation
Biology 34(1): 66–79. https://doi.org/10.1111/cobi.13308
Volis, S. (2019).
Conservation-oriented restoration – a two for one method to restore both
threatened species and their habitats. Plant Diversity 41: 50–58. https://doi.org/10.1016/j.pld.2019.01.002
Walpole, M.J. & H.J. Goodwin (2001). Local
attitudes towards conservation and tourism around Komodo National Park,
Indonesia. Environmental Conservation 28: 160–166. https://doi.org/10.1017/S0376892901000169
Whelan, C.J., D.G. Wenny & R.J. Marquis (2008). Ecosystem
services provided by birds. Annals of New York Academy of Sciences 1134:
25–60. https://doi.org/10.1196/annals.1439.003
Winter, S.J., K.J. Esler & M. Kidd (2005). An index to measure the conservation attitudes of landowners towards
Overberg Coastal Renosterveld, a critically endangered vegetation type in the
Cape Floral Kingdom, South Africa. Biological Conservation 126: 383–394. https://doi.org/10.1016/j.biocon.2005.06.015
WPA (1972). Wildlife
(Protection) Act. https://content.dgft.gov.in/Website/append10.pdf. Accessed on
10.xi.2025.
WPAA (2022). Wildlife
(Protection) Amendment Act. https://prsindia.org/files/bills_acts/acts_parliament/2022/The%20Wild%20Life%20(Protection)%20Amendment%20Act,%202022.pdf.
Accessed on 10.xi.2025.
Yashmita-Ulman &
M. Singh (2021). Bird composition, diversity and foraging guilds in agricultural
landscapes: A case study from eastern Uttar Pradesh, India. Journal of Threatened Taxa 13(8):
19011–19028. https://doi.org/10.11609/jott.7089.13.8.19011-19028
Yashmita-Ulman &
M. Singh (2022). Avifaunal diversity in unprotected wetlands of Ayodhya
district, Uttar Pradesh, India. Journal of Threatened Taxa 14(8):
21561–21578. https://doi.org/10.11609/jott.7067.14.8.21561-21578
Yashmita-Ulman &
M. Singh (2025). A preliminary assessment of avifaunal diversity in Parwati
Arga Bird Sanctuary, Uttar Pradesh, India. Journal
of Threatened Taxa 17(11): 27976–27984. https://doi.org/10.11609/jott.10160.17.11.27976-27984
Yashmita-Ulman
(2022). Bird diversity in riverscapes of Ayodhya
district, Uttar Pradesh. Indian
Journal of Ecology 49(1): 280–287. https://doi.org/10.55362/IJE/2022/3518
Yashmita-Ulman
(2023). Bird diversity of Jagdishpur Jheel: An unprotected wetland in Ayodhya
district, Uttar Pradesh. Indian Journal of Ecology 50(5) (S1):
1794–1800. https://doi.org/10.55362/IJE/2023/4139
Yashmita-Ulman, A. Kumar
& M. Sharma (2017). Traditional homegarden agroforestry systems: habitat for
conservation of Baya Weaver Ploceus
philippinus (Passeriformes: Ploceidae)
in Assam, India. Journal of Threatened Taxa 9(4): 10076–10083. https://doi.org/10.11609/jott.3090.9.4.10076-10083
Yashmita-Ulman, A.
Kumar & M. Sharma (2018). Agroforestry systems as habitat for avian species: Assessing its
role in conservation. Proceedings of Zoological Society 71: 127–145. https://doi.org/10.1007/s12595-016-0198-3
Yashmita-Ulman, M.
Singh, A. Kumar & M. Sharma (2020). Negative
human-wildlife interactions in traditional agroforestry systems in Assam,
India. Journal of Threatened Taxa 12(10): 16230–16238. https://doi.org/10.11609/jott.5754.12.10.16230-16238
Yashmita-Ulman, M.
Singh, A. Kumar & M. Sharma (2021a).
Agroforestry systems: a boon or bane
for mammal conservation in northeastern India? Proceedings of Zoological
Society 74: 28–42. https://doi.org/10.1007/s12595-020-00335-5
Yashmita-Ulman, M.
Singh, A. Kumar & M. Sharma (2021b).
Conservation of wildlife diversity in
agroforestry systems in eastern Himalayan biodiversity hotspot. Proceedings
of Zoological Society 74: 171–188. https://doi.org/10.1007/s12595-021-00361-x
Zhong, L.,
T. Li, Y. Li, T. Zou, T. Yu & C. Dai (2023). Local
farmers’ perceptions of ecosystem services and disservices provided by the
Black-necked Crane (Grus nigricollis) and
their conservation implications. Global Ecology and Conservation, 46:
e02614. https://doi.org/10.1016/j.gecco.2023.e02614