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Weather-induced mass deaths of the Common House Swift Apus affinis in ThengumarahadaVillage of the Nilgiris, southern India
B. Ramakrishnan 1, G. Sivasubramanian 2, K. Ramkumar 3 & S. Ramasubramanian 4
1 Assistant Professor of
Wildlife Biology, Department of Zoology & Wildlife Biology, Government Arts
College, Udhagamandalam, Tamil Nadu 643002 India
2 Senior Project Officer, Nilgiris & Eastern Ghats Landscape Programme,
WWF-India, Coimbatore, Tamil Nadu, India
3 Senior Field Officer, Wildlife Trust of
India, Noida 201301 India
4 District Forest Officer, Sathyamangalam Forest Division, Gobi Road, Sathyamangalam Post, Erode District, Tamil Nadu 638401
India
1 bio.bramki@gmail.com
(corresponding author), 2 sisu.wwf@gmail.com,3 ramkumar@wti.org.in, 4 uma_rama@mailcity.com
Abstract: Weather
influences all living organisms, especially wild animals. In the case of birds,
inclement weather conditions may affect them during migration and feeding that
could cause mass mortality in the same species or many related species. This paper describes an incidence of
mass deaths of the Common House Swift Apus affinis at ThengumarahadaVillage of Nilgiris, Tamil Nadu, southernIndia during February 2008. The
samples of the dead birds were sent for toxicological and bacteriological
examination at Southern Regional Disease Diagnostic Laboratory, Bangalore. Spot post mortems were also conducted by
forest veterinarians. Results of
the diagnostic analysis revealed that mass mortality might be due to extreme
weather conditions leading to acute dehydration, shock and death. The postmortemreports say that the crop and the gastro intestinal tract (GI) were found empty
in all the cases, suggesting starvation. We thus conclude that unseasonal heavy rains and non
availability of food could have led to hypothermia and starvation of
birds over a long period resulting in mass mortality.
Keywords:Apus affinis,
hypothermia, mass death, starvation, weather.
doi: http://dx.doi.org/10.11609/JoTT.o3405.5273-6 | ZooBank:urn:lsid:zoobank.org:pub:907FA4EF-E8DC-4C68-83BD-35FD9D42F86C
Editor: V.Santharam, Institute of Bird Studies & Natural
History, Chittoor, India. Date
of publication: 26 December 2013 (online & print)
Manuscript details: Ms #
o3405 | Received 10 November 2012 | Final received 02 December 2013 | Finally
accepted 04 December 2013
Citation: Ramakrishnan, B., G. Sivasubramanian, K. Ramkumar & S. Ramasubramanian (2013). Weather-induced mass deaths of the Common House Swift Apus affinis in Thengumarahada Village of the Nilgiris,
southern India. Journal
of Threatened Taxa 5(17): 5273–5276; http://dx.doi.org/10.11609/JoTT.o3405.5273-6
Copyright: © Ramakrishnan et al. 2013. Creative Commons Attribution 3.0 UnportedLicense. JoTT allows unrestricted use of this
article in any medium, reproduction and distribution by providing adequate
credit to the authors and the source of publication.
Funding: None.
Competing Interest: None.
Acknowledgements: We are deeply
indebted to the veterinary experts Dr. N.S. Manoharan, Forest Veterinary Officer, Dr.N. Kalaivanan, Forest Veterinary Assistant Surgeon,
Tamil Nadu Forest Department, Dr. R. Chandrasekaran, Assistant Director, Dr. Kasthuri and Dr. Yuvaraj, Veterinary Assistant Surgeons, Animal Disease
Intelligence Unit, Udhagamandalam for their kind help in sharing the post
mortem reports and laboratory results of the dead birds. We are thankful to Mr. G. Kannan, Biologist, Indian
Institute of Science, Masinagudi for providing
photographs (3 & 4) for this article. We are thankful to Mr. Soundrapandian, IFS, District
Forest Officer, Nilgiris North Forest Division, for
rendering his cooperation during the field observation. Our sincere thanks to Dr. C. Arivazhagan, Conservation
Biologist, Care Earth for his immense help in various
ways to complete this manuscript.
The publication of this article is
supported by the Critical Ecosystem Partnership Fund (CEPF), a joint initiative
of l’Agence Française de Développement, Conservation International, the
European Commission, the Global Environment Facility, the Government of
Japan, the MacArthur Foundation and the World Bank.
For figures, images -- click here
Climate change in response to global warming is one of the determining
factors in the conservation of wildlife in the present scenario. Living organisms occupy niche habitats
within a major ecosystem according to their biological requirements. The micro climatic conditions prevailing
within the ecological niche are responsible for the survival of living
organisms. Any abnormal variation
in the weather condition is likely to affect the survival of the native species
in their habitats. In the case of
birds, inclement weather conditions can affect the feeding activities and can
cause mass mortality owing to starvation (Newton 2007). Especially, for insectivorous bird
species, a sufficient quantity and accessibility of food is very crucial, which
in turn depends on favorable climatic conditions (Rajchard et al 2006). Insectivorous birds, especially swifts and similar aerial feeding birds
are extremely sensitive to weather conditions (Martins & Wright 1993). There were many reports on such
weather-induced mass bird deaths around the world (see http://home.att.net/~thehessians/birds.html).
This paper describes one such phenomenon of mass deaths of the Common
House Swift Apus affinisin Thengumarahada Village of the NilgirisDistrict of Tamil Nadu in southern India. The Common House Swift is a smoky-black bird with a prominent white
throat and rump and belongs to the order Apodiformes. It is a common breeding resident and has
a wide range of distribution, often associated with human habitations and
usually shelters in old buildings, forts, etc (Ali
& Ripley 1987). It has also
been listed as ‘Least Concern’ in the Threatened Birds of the World (Birdlife
International 2008; IUCN 2007). Swifts are generally seen in large flocks spending most of their time
feeding on insects while in flight and take rest only at night.
Study area
The Thengumrahada Village lies between 76054’34”–76056’0.8”N & 11033’19”–11034’40”E
and is situated amidst Nilgiri Eastern Slope Reserve
Forest (Fig. 1). About 500 acres of
land in this village was leased out to 141 families of the Badagacommunity, people of the Nilgiris, to cultivate
grains under “grow more food” campaign in the early 1950s. Crops such as Rice Oryza sativa, Maize Zeamays, Ragi Elusine coracana and Groundnut Arachishypogea are being cultivated in this village every year. Many insectivorous birds are attracted
every year towards the agricultural lands of this village due to huge
congregations of insects during the harvest period. Normally, paddy is harvested between
January and February when there is less chance of heavy rainfall.
Methods
From 12 to 14 February 2008, we recorded mass deaths of Apus affinis (1000+
dead birds) in and around Thengumarahada Village
immediately after heavy continuous rains. Pooled samples of the dead birds were sent to the Institute of Animal
Health and Veterinary Biologicals, Southern Regional
Disease Diagnostic Laboratory, Bangalore (SRDDL) on 15 February 2008 for
toxicological and bacteriological examinations. Spot post mortems were also conducted
jointly by the forest, local and district animal disease intelligence unit veterinarians. Rainfall data was collected from
2001–2008 for Thengumarahada Village from
Public Works Department, Government of Tamil Nadu.
Results
The final diagnostic result from SRDDL was received on 22 February 2008
stating that the tissue samples (lung/liver/heart/others) did not reveal growth
of any specific etiological pathogens, that the pooled samples of intestinal,
stomach contents, liver and kidneys were found negative for hydrocynide/zinc
phosphide/organo phosphorus compounds (OPC). The laboratory diagnosis result finally
concluded that the mass mortality might be due to extreme weather conditions
leading to acute dehydration, shock and death (for details see Appendix 1).
Since the laboratory results were negative, we switched focus to the
spot postmortem examinations. The postmortemresults revealed that the crop and the gastro-intestinal tracts (GI) were empty
in all cases, suggesting starvation. Besides, congestion of liver and kidney, viscous blood, pale and dry
mucous membrane and loss of fat reserves were also noticed (Images
1–4). The plumage of the
birds was completely wet and intertwined with mud. The above-mentioned observations
revealed that death was caused due to starvation and stress. The chicks and juveniles had not fed for
three days due to unavailability of food caused by heavy rains.
Within three days (12–14 Feburay 2008),
about 200mm rainfall was recorded which is quite unusual for this area compared
to the past few years (2001–2007) (Fig. 2). Flash floods were noticed in the nearby
jungle streams. The dead birds were
seen floating and the feathers were coated with muddy water. Most of the dead birds (nearly 80%) were
chicks and juveniles.
Discussion
ThengumarahadaVillage is in the rain shadow area which receives
rainfall from the northeast monsoon (September to November). From the records of the past few years
it is clear that this area receives rain from September to November. The unusual rainfall noticed in the
month of February 2008, may be the cause for the mass deaths of the
species. Food being scarce in the
forest, they would move to near-by agricultural fields for foraging. The Common House Swift is an aerial
feeder, predominantly dependent on winged insects. Owing to the heavy rains, insects may
have became dormant and the birds were without food
for three days. We did not find a
single dead specimen in the natural forest areas. Our observation continued for three
days from 12–14 February 2008. This present observation corroborates with previous sporadic incidents
that occurred around the world. Bull & Dawson (1969) stated that a total 810 birds belonging to 13
species were found dead during a three-day visit in South Canterbury due to a
severe and unseasonal snow-storm. They also recorded an absence of dead
birds in native forest areas. Avery
et al. (1977) found that cloud cover and wind directions were the reasons for
the mortality of many nocturnal birds in a North Dakota Tower, he recorded that
48 birds were found dead on 4 and 5 October 1972 due to overcast skies, and northeastern wind of 5–15 km/hr,
followed by 69 birds found dead due to overcast skies, and light rain on 25 and
26 September 1973.
This present observation indicates that heavy rain and non availability of food could have led to the starvation of
the birds. Due to starvation they
were not able to fly in the rain. Subsequently, these birds drowned as the storm water added more
weight. Also the heavy rain
saturated the plumage and increased the wing weight causing a loss of body heat
(Newton 2007). After the rain, some of the live birds were collected by the study team and
their wings were dried and they were allowed to fly. Most of them took off and continued to
survive after getting the necessary warmth.
The present observation concluded that the mass death of birds was due
to hypothermia and starvation. This
incident sheds light on the need for detailed studies of such incidents in the
light of climate change effects.
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