Bhutan Asiabell Codonopsis bhutanica Ludlow (Asterales: campanulaceae): a new addition to the Indian flora

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Samiran Panday
Vikash Kumar
Sudhansu Sekhar Dash
Bipin Kumar Sinha
Paramjit singh


Codonopsis bhutanica Ludlow, a lesser-known endemic species to central and eastern Himalaya, collected for the first time from Arunachal Pradesh, India.  The species was previously known only from Bhutan and Nepal.  The species is reported here with a detailed taxonomic description along with habitat image, a photo-plate illustrating different features and a map showing the locality in Arunachal Pradesh as a new record to Indian flora.


Article Details

Author Biographies

Samiran Panday, Budge Budge College, 7-D.C. Das Road, Shyampur, Kolkata, West Bengal 700137, India.

Senior Reseach Fellow

Vikash Kumar, Central National Herbarium, Botanical Survey of India, Howrah, West Bengal 711103, India.

Reserach Associates 

Bipin Kumar Sinha, Botanical Survey of India, CGO Complex, Sector 1, Salt Lake, Kolkata, West Bengal 700064, India.

Scientist F 

Paramjit singh, Botanical Survey of India, CGO Complex, Sector 1, Salt Lake, Kolkata, West Bengal 700064, India.

Forme Director

Botanical Surve of India 


Clarke, C.B. (1881). Campanulaceae, pp. 421–442. In: Hooker, J.D. (ed.). The Flora of British India, Vol. 3. L. Reeve & Co., London, 712pp.

Clement, R.A. (2001). Campanulaceae, pp. 1375–1397. In: Grierson, A.J.C. & D.G. Long (eds.). Flora of Bhutan, Vol. 2(3). Royal Botanic Garden, Edinburgh and Royal Government of Bhutan, 1675pp.

Dash, S.S. (2018). On Appraisal of the genus Codonopsis Wall. [Campanulaceae] in India. Abstracts of International Symposium on conservation of Angisperm Diverisity: Hidden Treasure of Today and Tomorrow. Dept. of Botany, The Maharaja Siyajirao Univeristy of Boroda, Vododara.

Dash, S.S. & A.A. Mao (2011). A new species of Codonopsis Wall. (Campanulaceae) from Arunachal Pradesh, India. Nelumbo 53: 17–20.

Giri, G.S., A. Pramanik & H.J. Chowdhery (2008). Materials for the Flora of Arunachal Pradesh, Vol. 2. Botanical Survey of India, Kolkata, 492pp.

Haridasan, V.K. & P.K. Mukherjee (1996). Campanulaceae, pp. 25–118. In: Hajra, P.K. & M. Sanjappa (eds.). Fascicles of Flora of India - Vol. 22. Botanical Survey of India, Calcutta, 144pp.

Hong, D. (2015). A Monograph of Codonopsis and Allied Genera (Campanulaceae). Science Press, Beijing, 270pp.

Hong, D. (2015). Campanulaceae, pp. 100–152. In: Flora of Pan-Himalaya. Vol. 47. Science Press, Beijing, 292pp.

Hong, D., G. Song, T.G. Lammers & L.L. Klein (2011). Campanulaceae, pp. 505–563. In: Wu, Z.G., P. Raven & D.Y. Hong (eds.). Flora of China. Vol. 19. Science Press, Beijing & Missouri Botanic Garden Press, St. Louis, USA.

Kanjilal, U.N., P.C. Kanjilal & A. Das (1939). Flora of Assam, Vol. 3. Govt. Press, Shillong, 588pp.

Kanwal, K.S., U.L. Tiwari, L. Yama & M.S. Lodhi (2019). Extended distribution record of two bellflower species of Codonopsis (Campanulaceae) from the Indian state of Arunachal Pradesh. Journal of Threatened Taxa 11(9): 14228–14231.

Mabberley, D.J. (2018). Mabberley’s Plant-Book: A Portable Dictionary of Plants, their Classifications and Uses, 4th Edition. Cambridge University Press, London, 215pp.

Mao, A.A. & H.A. Barbhuiya (2014). Extended distribution of Codonopsis tubulosa (Campanulaceae) in India. Nelumbo 56: 234–237.

Panday, S. & B.K. Sinha (2012). Rediscovery of Campanumoea javanica Blume- a rare climber of Campanulaceae from Phawngpui National Park in Mizoram, India. Pleione 6(1): 241–243.